Udelartitan

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Udelartitan
Temporal range: Upper Cretaceous
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Clade: Macronaria
Clade: Titanosauria
Superfamily: Saltasauroidea
Family: incertae sedis
Genus: Udelartitan
Species:
U. celeste
Binomial name
Udelartitan celeste
Soto et al., 2024

Udelartitan is an extinct genus of saltasauroid titanosaurian sauropod dinosaur from the Late Cretaceous Guichón Formation of Uruguay. The genus contains a single species, U. celeste, known from fragmentary remains of at least two individuals.

Discovery and naming

The Udelartitan fossil material was discovered in 2006 within sediments of the Guichón Formation (Araújo locality), near Quebracho in Paysandú Department, Uruguay. The holotype specimen, FC-DPV 3595, consists of the first three caudal vertebrae in approximate articulation. Additional bones representing an older individual (specimen FC-DPV 1900), including sixty caudal vertebrae, an incomplete left coracoid, the proximal and distal ends of the tibiae the proximal end of the fibula, six metatarsals, and two astragali, were also referred to Udelartitan. [1] Many of the bones of the referred specimen were preliminarily described by Soto, Perea & Cambiaso in 2012, who also identified several purported osteoderms. [2] However, Soto et al. (2024) found no evidence to support this assessment, arguing that they can more confidently be described as concretions. [1]

In 2024, Soto et al. described Udelartitan celeste as a new genus and species of titanosaur sauropod based on these fossil remains. The generic name, Udelartitan, combines "UdelaR", an acronym for the Universidad de la República (whose collection of vertebrae fossils includes those assigned to Udelartitan), with the word "titan", a common suffix for giant sauropod names, referencing the pre-Olympian gods of Greek mythology. The specific name, celeste, is a Spanish word for sky blue, referencing the nickname given to some Uruguayan athletic teams. [1]

Description

Based on other titanosaurs known from more complete skeletons, Soto et al. (2024) suggest that Udelartitan was about 15–16 metres (49–52 ft) in body length. [1]

Classification

In their preliminary description of some of the Udelartitan fossil material in 2012, Soto, Perea & Cambiaso used the general morphology of the caudal vertebrae and astragalus to suggest that it likely belonged to a derived lithostrotian titanosaur—likely a non-saltasaurine saltasaurid—related to Pellegrinisaurus , Baurutitan , and Alamosaurus . They further cautioned that the Saltasauridae clade has historically had a very variable composition, comprising either very few or many species, depending on the analyses performed. [2]

In 2024, Soto et al. used phylogenetic analyses to find the most likely position for Udelartitan within the Titanosauria. Similar to the previous work, they found support for a position within the Saltasauroidea. They used two data-matrices to assess the phylogenetic position of Udelartitan, and recovered it either as a non-saltasaurid saltasauroid or a basal saltasaurine. Their results are displayed in the cladograms below: [1]

Palaeoenvironment

Life restoration of the contemporary Uruguaysuchus Uruguaysuchus BW.jpg
Life restoration of the contemporary Uruguaysuchus

Udelartitan is known from the Late Cretaceous Guichón Formation of Uruguay. The exact age of this formation is uncertain. Some authors have historically suggested a Lower Cretaceous (AptianAlbian) age due to similarities with the Migues Formation. [5] However, the overlying Mercedes Formation dates to the CampanianMaastrichtian ages of the Upper Cretaceous, so this would introduce a significant gap in time. The discovery of dinosaur eggshells in the formation similar to Sphaerovum prompted later researchers to favor a Late Cretaceous age for the Guichón Formation. [2]

Other fossils from the Guichón Formation include several specimens of Uruguaysuchus —a crocodyliform closely related to Araripesuchus —as well as the teeth of indeterminate iguanodontians and theropods. [6] [2] The younger Asencio Formation of Uruguay also contains titanosaur specimens, including the centrum of a caudal vertebra referred to Aeolosaurus sp. [1]

Related Research Articles

<span class="mw-page-title-main">Titanosauria</span> Extinct clade of dinosaurs

Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan—estimated at 37 m (121 ft) long with a weight of 69 tonnes —and the comparably-sized Argentinosaurus and Puertasaurus from the same region.

<i>Alamosaurus</i> Extinct genus of dinosaurs

Alamosaurus is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to Argentinosaurus and Puertasaurus, which would make it the absolute largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age. Specimens of a juvenile Alamosaurus sanjuanensis have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Alamosaurus is the only known sauropod to have inhabited North America after their nearly 30-million year absence from the North American fossil record and probably represents an immigrant from South America.

<i>Bonitasaura</i> Extinct genus of dinosaurs

Bonitasaura is a genus of titanosaurian dinosaur hailing from uppermost layers of the Late Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén Group of the eastern Neuquén Basin, located in Río Negro Province, Northwestern Patagonia, Argentina. The remains, consisting of a partial sub-adult skeleton jumbled in a small area of fluvial sandstone, including a lower jaw with teeth, a partial vertebrae series, and limb bones, were described by Sebastian Apesteguía in 2004.

<i>Adamantisaurus</i> Extinct genus of dinosaurs

Adamantisaurus is a poorly-known genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. It is only known from six tail vertebrae but, as a sauropod, it can be assumed that this dinosaur was a very large animal with a long neck and tail.

<i>Epachthosaurus</i> Extinct genus of dinosaurs

Epachthosaurus was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America.

<i>Neuquensaurus</i> Extinct genus of dinosaurs

Neuquensaurus is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived.

<i>Baurutitan</i> Extinct genus of dinosaurs

Baurutitan is a genus of sauropod dinosaur that lived during the Late Cretaceous in what is now Brazil. The type species, Baurutitan britoi, was described in 2005 by Kellner and colleagues, although the fossil remains had already been discovered in 1957. Baurutitan is classified as a lithostrotian titanosaur, and is distinguished from related genera based on its distinctive caudal vertebrae. This South American dinosaur was found in the Serra da Galga Formation near Uberaba, in the Brazilian state of Minas Gerais.

<i>Rocasaurus</i> Genus of titanosaurian sauropod from the Late Cretaceous period

Rocasaurus is a genus of titanosaurian sauropod that lived in South America. Rocasaurus was discovered in Argentina in 2000, within the Allen Formation which is dated to be middle Campanian to early Maastrichtian in age. This genus grew up to 8 metres (26 ft) long, making it one of the smaller sauropods. It seems to be closely related to saltasaurid dinosaurs, like Saltasaurus and Neuquensaurus.

<i>Pellegrinisaurus</i> Extinct genus of dinosaurs

Pellegrinisaurus is a genus of titanosaurian sauropod dinosaur that lived in South America during the Late Cretaceous period. The holotype was found in the Allen Formation, Argentina.

<i>Puertasaurus</i> Extinct genus of dinosaurs

Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.

<span class="mw-page-title-main">Lithostrotia</span> Extinct clade of dinosaurs

Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.

Malarguesaurus is a genus of titanosauriform sauropod dinosaur from the Late Cretaceous of Mendoza Province, Argentina. Its fossils, consisting of tail vertebrae, chevrons, ribs, and limb bones, were found in the upper Turonian-lower Coniacian Portezuelo Formation of the Neuquén Group. The type species, described by González Riga et al. in 2008, is M. florenciae.

The Guichón Formation is a Late Cretaceous geologic formation of the Paysandú Group in Uruguay. Dinosaur remains are among the fossils that have been recovered from the formation.

Sphaerovum is an oogenus of dinosaur egg that has only been discovered in South America.

<span class="mw-page-title-main">Aeolosaurini</span> Extinct clade of dinosaurs

Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found Aeolosaurus, Gondwanatitan, Maxakalisaurus, Panamericansaurus and Rinconsaurus to be aeolosaurids.

Brasilotitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Adamantina Formation of Brazil. The type species is Brasilotitan nemophagus. Brasilotitan was a small titanosaur with a squared-off snout, and may be closely related to another Brazilian titanosaur, Uberabatitan.

<span class="mw-page-title-main">Saltasauroidea</span>

Saltasauroidea is a superfamily of titanosaurs named by França and colleagues in 2016 based on their phylogenetic results, for a clade uniting Aeolosaurini and Saltasauridae, as well as the intermediate genera Baurutitan, Diamantinasaurus and Isisaurus. The group was not defined or discussed in the text, but was supported by Carballido and colleagues in 2022 as a useful designation for subdividing titanosaurs. As there was no discussion about the intentions for the clade, Carballido gave it the definition of all taxa closer to Saltasaurus than Patagotitan, encompassing half of Eutitanosauria as the sister taxon to the inversely defined Colossosauria. Carballido et al. placed Nemegtosauridae and Saltasauridae within the group, though they had Aeolosaurini within Colossosauria. The informal cladogram of titanosaur relationships they proposed is shown below.

Ibirania is a genus of dwarf saltasaurine titanosaur dinosaur from the Late Cretaceous São José do Rio Preto Formation of Southeast Brazil. The type species is Ibirania parva. It is one of the smallest sauropods known to date, comparable in size to the titanosaur Magyarosaurus.

Sidersaura is an extinct genus of rebbachisaurid sauropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, S. marae, known from the remains of four individuals. Sidersaura represents one of the largest known rebbachisaurids.

<i>Chakisaurus</i> Extinct genus of ornithopod dinosaurs

Chakisaurus is an extinct genus of elasmarian ornithopod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, C. nekul, known from multiple partial skeletons belonging to individuals of different ages. Chakisaurus represents the first ornithischian species to be named from the Huincul Formation.

References

  1. 1 2 3 4 5 6 Soto, Matías; Carballido, José L.; Langer, Max C.; Silva Junior, Julian C. G.; Montenegro, Felipe; Perea, Daniel (2024). "Phylogenetic relationships of a new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Uruguay". Cretaceous Research . 160. 105894. doi:10.1016/j.cretres.2024.105894.
  2. 1 2 3 4 Soto, Matías; Perea, Daniel; Cambiaso, Andrea (2012-02-01). "First sauropod (Dinosauria: Saurischia) remains from the Guichón Formation, Late Cretaceous of Uruguay". Journal of South American Earth Sciences . 33 (1): 68–79. Bibcode:2012JSAES..33...68S. doi:10.1016/j.jsames.2011.08.003. ISSN   0895-9811.
  3. Cerda, Ignacio; Zurriaguz, Virginia Laura; Carballido, José Luis; González, Romina; Salgado, Leonardo (2021). "Osteology, paleohistology and phylogenetic relationships of Pellegrinisaurus powelli (Dinosauria: Sauropoda) from the Upper Cretaceous of Argentinean Patagonia". Cretaceous Research . 128: 104957. Bibcode:2021CrRes.12804957C. doi:10.1016/j.cretres.2021.104957.
  4. Navarro, Bruno A.; Ghilardi, Aline M.; Aureliano, Tito; Díaz, Verónica Díez; Bandeira, Kamila L. N.; Cattaruzzi, André G. S.; Iori, Fabiano V.; Martine, Ariel M.; Carvalho, Alberto B.; Anelli, Luiz E.; Fernandes, Marcelo A.; Zaher, Hussam (2022-09-15). "A new nanoid titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Brazil". Ameghiniana. 59 (5): 317–354. doi:10.5710/AMGH.25.08.2022.3477. ISSN   1851-8044. S2CID   251875979.
  5. de Santa Ana, Héctor; Goso Aguilar, César; Montaño, Jorge; Piñero, Graciela; Muzio, Rossana; Rossello, Eduardo; Perea, Daniel; Ucha, Nelson (2004). "El Cretácico post-basáltico y el Terciario Inferior de la Cuenca litoral del río Uruguay: geología y paleontología.". In Veroslavsky, Gerardo; Uvilla, Martín; Martínez, Sergio (eds.). Cuencas Sedimentarias de Uruguay: Geología, paleontología y recursos naturales (PDF) (in Spanish) (2nd ed.). Universidad de la República. pp. 143–171.
  6. Soto, Matías; Pol, Diego; Perea, Daniel (2012-01-12). "A new specimen of Uruguaysuchus aznarezi (Crocodyliformes: Notosuchia) from the middle Cretaceous of Uruguay and its phylogenetic relationships". Zoological Journal of the Linnean Society . 163: S173–S198. doi:10.1111/j.1096-3642.2011.00717.x. hdl: 11336/69086 .