Coloradisaurus | |
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Diagram showing known skull elements; shaded parts represent missing bones | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Family: | † Massospondylidae |
Genus: | † Coloradisaurus Galton, 1990 [2] [3] |
Type species | |
†Coloradisaurus brevis | |
Synonyms | |
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Coloradisaurus (meaning "Los Colorados lizard") is a genus of massospondylid sauropodomorph dinosaur. It lived during the Late Triassic period (Norian stage) in what is now La Rioja Province, Argentina. It is known from two specimens collected from the Los Colorados Formation of the Ischigualasto-Villa Unión Basin.
Coloradisaurus brevis was originally named Coloradia brevis by José Bonaparte in 1978, [4] but that genus name was preoccupied by the pine moth Coloradia , [5] so it needed a replacement name. In 1983, David Lambert used the name Coloradisaurus for the genus, but did not indicate it was a replacement or diagnose it. [6] Lambert had gotten the name from Bonaparte in a personal communication and mistakenly thought that Bonaparte had already published it. [3] Peter Galton was the next to use the name Coloradisaurus in 1990, which he credited to Lambert, when he gave the taxon a diagnosis in his review of prosauropods in The Dinosauria . [2] [3] Authorship of Coloradisaurus has traditionally been attributed to Lambert, but in 2020, Greenfield et al. judged Lambert's use of the name to be a nomen nudum . They concluded that authorship should be attributed to Galton, who was the first to use the name Coloradisaurus in a way that met the requirements of the ICZN. [3]
The holotype of Coloradisaurus (PVL 3967) is a mostly complete skull found associated with an undescribed partial skeleton. [4] While the right side of the skull is well-preserved with almost all bones intact, the left side is distorted and missing more bones. [7] The holotype individual has been estimated to have been 3 m (10 ft) long with a mass of 70 kg (150 lb). [8] A referred specimen (PVL 5904) is a partial skeleton including the most of the dorsal vertebrae and parts of the pectoral and pelvic girdles and limbs. [9] Like Lufengosaurus , it have the angle between the pterygoid and quadratojugal rami nearly 90°. There is also a possibility that the postorbital bones of Coloradisaurus and Sarahsaurus are similar, but due to the deformation of the skull this is difficult to say. [10]
All material of Coloradisaurus was discovered in 1971 at the La Esquina locality in the upper section of the Los Colorados Formation near Pagancillo, La Rioja Province, Argentina. [4] [7] [9] The top of the Los Colorados Formation has been dated to 213 Ma, [1] which would place Coloradisaurus in the Norian stage of the Late Triassic.
Coloradisaurus was classified as a plateosaurid in the original description by Bonaparte, [4] but this pre-dated the use of phylogenetic analyses in paleontology. He later became opposed to cladistics [11] and continued to consider Coloradisaurus a plateosaurid without testing its phylogenetic position. [12] The analyses of Galton (1990), Galton & Upchurch (2004), and Upchurch et al. (2007) found it to be a plateosaurid, supporting Bonaparte's placement. [2] [13] [14] However, the analyses of Benton et al. (2000) and Yates (2003) recovered it in a polytomy with other basal sauropodomorphs or as a massospondylid, respectively. [15] [16] Subsequent analyses such as Yates et al. (2010), Apaldetti et al. (2013; 2014), Wang et al. (2017), and Müller (2020) have reached the consensus that Coloradisaurus is a massospondylid most closely related to Lufengosaurus and Glacialisaurus . [7] [9] [17] [18] [10] [19] These three taxa share four synapomorphies found in the metatarsals and femur. [9]
Below is a simplified cladogram after Galton & Upchurch (2004), reflecting its early placement as a plateosaurid. [13]
Sauropodomorpha |
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Below is a simplified cladogram after Müller (2020), showing its current position as a massospondylid. [19]
Plateosauria |
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Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.
Saturnalia is an extinct genus of basal sauropodomorph dinosaur known from the Late Triassic Santa Maria Formation of Rio Grande do Sul, southern Brazil. It is one of the earliest known dinosaurs.
Herrerasaurus is likely a genus of saurischian dinosaur from the Late Triassic period. This genus was one of the earliest dinosaurs from the fossil record. Its name means "Herrera's lizard", after the rancher who discovered the first specimen in 1958 in South America. All known fossils of this carnivore have been discovered in the Ischigualasto Formation of Carnian age in northwestern Argentina. The type species, Herrerasaurus ischigualastensis, was described by Osvaldo Reig in 1963 and is the only species assigned to the genus. Ischisaurus and Frenguellisaurus are synonyms.
Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Africa, it had a large body and sturdy limbs, suggesting it moved about on all fours. Its limb bones were massive and heavy like the limb bones of true sauropods.
Riojasaurus was a herbivorous sauropodomorph dinosaur named after La Rioja Province in Argentina where it was found in the Los Colorados Formation in the Ischigualasto-Villa Unión Basin by José Bonaparte. It lived during the Late Triassic and grew to about 6.6 metres (22 ft) long. Riojasaurus is the only known riojasaurid to live in South America.
Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.
Lessemsaurus is an extinct genus of sauropod dinosaur belonging to Lessemsauridae.
Euskelosaurus is a sauropodomorph dinosaur from the Late Triassic of South Africa and Lesotho. Fossils have only been recovered from the lower Elliot Formation in South Africa and Lesotho, and in one locality in Zimbabwe.
Peter Malcolm Galton is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosauropod dinosaurs.
Plateosauridae is a family of plateosaurian sauropodomorphs from the Late Triassic of Europe, Greenland, Africa and Asia. Although several dinosaurs have been classified as plateosaurids over the years, the family Plateosauridae is now restricted to Plateosaurus, Yimenosaurus, Euskelosaurus, and Issi. In another study, Yates (2003) sunk Sellosaurus into Plateosaurus. Gresslyosaurus is alternatively considered its own genus or a synonym of Plateosaurus.
Guaibasauridae is a family of basal sauropodomorph dinosaurs, known from fossil remains of late Triassic period formations in Brazil and Argentina.
The Los Colorados Formation is a sedimentary rock formation of the Ischigualasto-Villa Unión Basin, found in the provinces of San Juan and La Rioja in Argentina. The formation dates back to the Norian age of the Late Triassic.
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.
Anchisauria is an extinct clade of sauropodomorph dinosaurs that lived from the Late Triassic to the Late Cretaceous. The name Anchisauria was first used Haekel and defined by Galton and Upchurch in the second edition of The Dinosauria. It is a node-based taxon containing the most recent common ancestor of Anchisaurus polyzelus and Melanorosaurus readi, and all its descendants. Galton and Upchurch assigned a family of dinosaurs to the Anchisauria: the Melanorosauridae. The more common prosauropods Plateosaurus and Massospondylus were placed in the sister clade Plateosauria.
Glacialisaurus is a genus of sauropodomorph dinosaur. It lived during the Pliensbachian stage of the Early Jurassic period around 186 to 182 million years ago in what is now the central region of the Transantarctic Mountains of Antarctica. It is known from two specimens; the holotype, a partial tarsus (ankle) and metatarsus, and a partial left femur. The fossils were collected by a team led by paleontologist William R. Hammer during a 1990–91 field expedition to the Hanson Formation of Antarctica. They were described in 2007, and made the basis of the new genus and species Glacialisaurus hammeri. The genus name translates as “icy” or "frozen lizard”, and the species name honors Hammer.
Plateosauria is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to the Late Cretaceous. The name Plateosauria was first coined by Gustav Tornier in 1913. The name afterwards fell out of use until the 1980s.
Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defined as all animals more closely related to Saltasaurus loricatus than to Plateosaurus engelhardti.
The Melanorosauridae were a family of sauropodomorph dinosaurs which lived during the Late Triassic and Early Jurassic. The name Melanorosauridae was first coined by Friedrich von Huene in 1929. Huene assigned several families of dinosaurs to the infraorder "Prosauropoda": the Anchisauridae, the Plateosauridae, the Thecodontosauridae, and the Melanorosauridae. Since then, these families have undergone numerous revisions. Galton and Upchurch (2004) considered Camelotia, Lessemsaurus, and Melanorosaurus members of the family Melanorosauridae. A more recent study by Yates (2007) indicates that the melanorosaurids were instead early sauropods.
Adeopapposaurus is a genus of plateosaurian dinosaur from the Early Jurassic Cañón del Colorado Formation of San Juan, Argentina. It was similar to Massospondylus. Four partial skeletons with two partial skulls are known.
Xingxiulong is a genus of bipedal sauropodiform from the Early Jurassic of China. It contains a single species, X. chengi, described by Wang et al. in 2017 from three specimens, two adults and an immature individual, that collectively constitute a mostly complete skeleton. Adults of the genus measured 4–5 metres (13–16 ft) long and 1–1.5 metres tall. Phylogenetic analysis suggests that Xingxiulong is most closely related to its contemporary Jingshanosaurus, although an alternative position outside of both the Sauropodiformes and Massospondylidae is also plausible.
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