Gondwanatitan Temporal range: Late Cretaceous, | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Clade: | † Lithostrotia |
Genus: | † Gondwanatitan Kellner & de Azevedo, 1999 |
Species: | †G. faustoi |
Binomial name | |
†Gondwanatitan faustoi Kellner & de Azevedo, 1999 | |
Gondwanatitan (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. Gondwanatitan was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other sauropods, Gondwanatitan was tall and ate tough shoots and leaves off of the tops of trees. G. faustoi's closest relative was Aeolosaurus .
The type species is Gondwanatitan faustoi, formally described by Kellner and de Azevedo in 1999.
Gondwanatitan means "Gondwana Titan", and is named after Gondwana, the supercontinent that the genus' South American range was once part of, and the Titans of classical Greek mythology. The type and only named species, G. faustoi, is a patronym honoring Dr. Fausto L. de Souza Cunha, a former curator at the Museu Nacional/UFRJ who led the excavation of the type specimen. [1]
Gondwanatitan was a fairly small sauropod, only 7 meters (23 ft) long and weighing about 1 tonne (1.1 short tons). [2] It had relatively gracile limb bones. [1] The middle caudal vertebrae are distinctively "heart-shaped", which allows isolated caudal vertebrae to be easily distinguished from those of Aeolosaurus. [3]
The vertebrae from the middle part of its tail had elongated centra. [4] Gondwanatitan had vertebral lateral fossae that resembled shallow depressions. [4] Fossae that similarly resemble shallow depressions are known from Saltasaurus , Alamosaurus , Malawisaurus , and Aeolosaurus . [4] Its middle tail vertebrae's neural spines are angled anteriorly when the vertebrae are aligned. [4] These vertebrae resemble those of Cedarosaurus , Venenosaurus , and Aeolosaurus . [4]
Gondwanatitan is a member of the clade Aeolosauridae. [5] It is closely related to the genera Pitekunsaurus , Aeolosaurus , and Overosaurus . [6]
The type specimen of Gondwanatitan faustoi was found in strata of the Adamantina Formation. [1] Other material assigned to the genus has been found in the Cambabe Formation. [5]
The type specimen of Gondwanatitan faustoi was discovered in 1983 on the farm of Yoshitoshi Myzobuchi in São Paulo, Brazil. [1] The specimen was excavated between 1984 and 1986, but preparation work on the specimen did not begin in earnest until 1997. It was finally described as a new genus and species in 1999. In 2001, G. faustoi was briefly transferred to the genus Aeolosaurus, making Gondwanatitan a junior synonym of that genus, but it has since been widely regarded as separate. [7] [8]
Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan—estimated at 37 m (121 ft) long with a weight of 69 tonnes —and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species A. maximus was transferred over to the new genus Arrudatitan in 2021.
Malawisaurus is an extinct genus of titanosaurian sauropod dinosaur. It is known from the Dinosaur Beds of northern Malawi, which probably date to the Aptian stage of the Early Cretaceous. The type species is M. dixeyi and the specific name honours Frederick Augustus Dixey.
Adamantisaurus is a poorly-known genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. It is only known from six tail vertebrae but, as a sauropod, it can be assumed that this dinosaur was a very large animal with a long neck and tail.
Cedarosaurus was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period (Valanginian). It was a sauropod which lived in what is now Utah. The fossils were discovered in 1996 in eastern Utah within the rocks of the Yellow Cat Member of the Cedar Mountain Formation. It was officially named and described by Tidwell, Carpenter and Brooks in 1999.
Venenosaurus is a genus of sauropod dinosaur that lived in what is now Utah during the Early Cretaceous. Its type and only species is Venenosaurus dicrocei. Fossils of Venenosaurus were first discovered in 1998, by Denver Museum of Natural History volunteer Anthony DiCroce, and described as a new genus and species in 2001 by Virginia Tidwell and colleagues, who named the species for DiCroce. Venenosaurus was a relatively small sauropod, and was similar to Cedarosaurus, another sauropod from the Early Cretaceous of Utah.
Rinconsaurus is a genus of titanosaur sauropod dinosaur from the Late Cretaceous in what is now Argentina. The type species, Rinconsaurus caudamirus, was described by Calvo and Riga in 2003, and is based on three partial skeletons.
Maxakalisaurus is a genus of titanosaur dinosaur, found in the Adamantina Formation of Brazil, in the state of Minas Gerais in 1998. The genus name is derived from the tribe of the Maxakali; Topa is one of their divinities.
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.
Uberabatitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of Brazil. It is known from bones including neck, back, and tail vertebrae, pelvic bones, and limb bones. These fossils were found in the uppermost portion of the Maastrichtian-age Serra da Galga Formation of the Bauru Group, in Uberaba, Minas Gerais. The type species, described by Salgado and Carvalho in 2008, is U. ribeiroi. To date, it is the most recent titanosaur from Bauru Group rocks; other titanosaurs from the Bauru Group, including Baurutitan and Trigonosaurus, come from lower levels.
Pitekunsaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Anacleto Formation of Neuquén, Argentina. It was described by L. Filippi and A. Garrido in 2008. The type species is P. macayai. The generic name is derived from Mapudungun pitekun, meaning "to discover", the epitheton honours the discoverer, oil company explorer Luis Macaya, who found the fossil in April 2004.
The Adamantina Formation is a geological formation in the Bauru Basin of western São Paulo state, in southeastern Brazil.
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found Aeolosaurus, Gondwanatitan, Maxakalisaurus, Panamericansaurus and Rinconsaurus to be aeolosaurids.
Overosaurus is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia. Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft). It was a ground-dwelling herbivore.
Brasilotitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Adamantina Formation of Brazil. The type species is Brasilotitan nemophagus. Brasilotitan was a small titanosaur with a squared-off snout, and may be closely related to another Brazilian titanosaur, Uberabatitan.
Austroposeidon is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Presidente Prudente Formation of Brazil. It contains one species, Austroposeidon magnificus.
Rinconsauria is an extinct clade of giant titanosaurian sauropods known from the late Cretaceous period of Argentina.
Arrudatitan is an extinct genus of titanosaur sauropod dinosaur known from the Late Cretaceous (Campanian-Maastrichtian)-aged Adamantina Formation of Brazil. The type species, A. maximus, was named and described in 2011 as a species of Aeolosaurus, but was separated into its own genus in 2021. It was relatively gracile for a titanosaur.
Ibirania is a genus of dwarf saltasaurine titanosaur dinosaur from the Late Cretaceous São José do Rio Preto Formation of Southeast Brazil. The type species is Ibirania parva. It is one of the smallest sauropods known to date, comparable in size to the titanosaur Magyarosaurus.
Caieiria is a genus of titanosaurian dinosaur from the Late Cretaceous Serra da Galga Formation of Brazil. The type and only species is Caieiria allocaudata.
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