Bustingorrytitan Temporal range: Late Cretaceous, | |
---|---|
Forelimb bones | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Clade: | † Lithostrotia |
Genus: | † Bustingorrytitan Simón & Salgado, 2023 |
Species: | †B. shiva |
Binomial name | |
†Bustingorrytitan shiva Simón & Salgado, 2023 | |
Bustingorrytitan (meaning "Bustingorry's giant") is a genus of lithostrotian titanosaur from the Late Cretaceous (Cenomanian) Huincul Formation of Argentina. The type species is Bustingorrytitan shiva. [1]
In 2001, several sauropod skeletons were uncovered at the "Bustingorry II" site at Villa El Chocón, including remains belonging to both Choconsaurus and Bustingorrytitan. [2] This sauropod was announced in a conference the same year by Simón, and received the informal name of "Sauropodus". [3] It was first described in 2011, in a thesis by María Edith Simón. [4]
It was named as a new genus and species of titanosaur in 2023. The generic name, "Bustingorrytitan", honors Manuel Bustingorry, the person who owns the land where the fossils were found, combining his name with the Greek titan, in reference to its large size. The specific name, "shiva", is named after the Hindu god Shiva, who destroys and transforms the universe, in reference to the Cenomanian-Turonian faunal turnover. [1]
The holotype of Bustingorrytitan is MMCH-Pv 59/1-40, a partial skeleton including a tooth, vertebra, possible ribs, a scapula, a humerus, a femur, tibiae and a fibula. Four specimens are known, together including parts of the dentary and postcranial skeleton. [1]
Using femur allometry, the weight of Bustingorrytitan was estimated to be approximately 67.3 tonnes (66.2 long tons; 74.2 short tons), making it one of the largest titanosaurs. It is also likely that the specimen doesn't belong to a fully grown animal. [1] However, estimates based on limb bone allometry have been criticized as unreliable, and when applied to large sauropods regularly produce higher mass estimates than those produced by other methods. [5] [6] [7]
Bustingorrytitan was entered into a phylogenetic analysis and recovered as a member of the Lithostrotia, which itself is recovered as a subclade of the Saltasauroidea, in contrast to most analyses. The results of the analysis are shown in the cladogram below: [1]
Bustingorrytitan was recovered from the Huincul Formation alongside two other gigantic titanosaurs, Argentinosaurus and Chucarosaurus . The three are not closely related to one another, which means that giant titanosaurs evolved multiple times within Eutitanosauria. [1]
Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, measuring 30–35 m (98–115 ft) long and weighing 65–80 t. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.
Antarctosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. The type species, Antarctosaurus wichmannianus, and a second species, Antarctosaurus giganteus, were described by prolific German paleontologist Friedrich von Huene in 1929. Three additional species of Antarctosaurus have been named since then but later studies have considered them dubious or unlikely to pertain to the genus.
Limaysaurus is a genus represented by a single species of rebbachisaurid sauropod dinosaurs, which lived during the mid-Cretaceous period, about 99.6 to 97 million years ago, in the Cenomanian, in what is now South America.
Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.
The Candeleros Formation is a geologic formation that crops out in the Río Negro, Neuquén, and Mendoza provinces of northern Patagonia, Argentina. It is the oldest formation in the Neuquén Group and belongs to the Rio Limay Subgroup. Formerly that subgroup was treated as a formation, and the Candeleros Formation was known as the Candeleros Member.
The Huincul Formation is a geologic formation of Late Cretaceous age of the Neuquén Basin that outcrops in the Mendoza, Río Negro and Neuquén Provinces of northern Patagonia, Argentina. It is the second formation in the Río Limay Subgroup, the oldest subgroup within the Neuquén Group. Formerly that subgroup was treated as a formation, and the Huincul Formation was known as the Huincul Member.
Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.
Skorpiovenator is a genus of abelisaurid theropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. It is one of the most complete and informative abelisaurids yet known, described from a nearly complete and articulated skeleton.
Patagotitan is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from at least six young adult individuals, Patagotitan mayorum, which was first announced in 2014 and then named in 2017 by José Carballido and colleagues. Preliminary studies and press releases suggested that Patagotitan was the largest known titanosaur and land animal overall, with an estimated length of 37 m (121 ft) and an estimated weight of 69 tonnes. Later research revised the length estimate down to 31 m (102 ft) and weight estimates down to approximately 50–57 tonnes, suggesting that Patagotitan was of a similar size to, if not smaller than, its closest relatives Argentinosaurus and Puertasaurus. Still, Patagotitan is one of the most-known titanosaurs, and so its interrelationships with other titanosaurs have been relatively consistent in phylogenetic analyses. This led to its use in a re-definition of the group Colossosauria by Carballido and colleagues in 2022.
Notocolossus is a genus of titanosaurian sauropod dinosaur from late Cretaceous strata of Mendoza Province, Argentina.
Taurovenator is a large carcharodontosaurid theropod from the late Cretaceous Huincul Formation of Argentina that lived during the Cenomanian age of the Late Cretaceous. It is monotypic, containing only one species, T. violantei.
Choconsaurus is an extinct genus of herbivorous sauropod dinosaur belonging to the group Titanosauriformes, which lived in the area of present-day Argentina at the end of the Cretaceous.
The Ernesto Bachmann Paleontological Museum (MEB) in Villa El Chocón, Neuquén Province, Argentina, is a municipal museum dedicated to the paleontology, archaeology and history of Villa El Chocón and its surroundings.
Kaikaifilusaurus is an extinct genus of rhynchocephalians in the family Sphenodontidae from the Late Cretaceous of South America. Fossils of the genus were found in Cenomanian sediments of the Candeleros Formation and Turonian layers of the Huincul Formation, both of the Neuquén Basin and the Albian strata of the Cerro Barcino Formation in the Cañadón Asfalto Basin, all in Patagonia, Argentina. The genus contains two species, K. minimus and the type species K. calvoi.
Overoraptor is an extinct genus of paravian theropod of uncertain affinities from the Late Cretaceous Huincul Formation of Argentinian Patagonia. The genus contains a single species, O. chimentoi, known from several bones of the hands, feet, and hips alongside some vertebrae.
Meraxes is a genus of large carcharodontosaurid theropod dinosaur from the Late Cretaceous Huincul Formation of Patagonia, Argentina. The genus contains a single species, Meraxes gigas.
Chucarosaurus is an extinct genus of titanosaurian dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, C. diripienda, known from various limb and pelvic bones.
Sidersaura is an extinct genus of rebbachisaurid sauropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, S. marae, known from the remains of four individuals. Sidersaura represents one of the largest known rebbachisaurids.
Chakisaurus is an extinct genus of elasmarian ornithopod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, C. nekul, known from multiple partial skeletons belonging to individuals of different ages. Chakisaurus represents the first ornithischian species to be named from the Huincul Formation.
Campananeyen is a genus of rebbachisaurid sauropod dinosaur from the Late Cretaceous (Cenomanian) Candeleros Formation of Argentina. The genus contains a single species, Campananeyen fragilissimus, known from a fragmentary skeleton.