Cardiodon Temporal range: late Bathonian ~ | |
---|---|
Holotype tooth seen from two different angles (2–3), another tooth (4) and a close-up of the surface of the tooth (5) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Turiasauria |
Genus: | † Cardiodon Owen, 1841 |
Species: | †C. rugulosus |
Binomial name | |
†Cardiodon rugulosus Owen, 1844 | |
Synonyms | |
|
Cardiodon (meaning "heart tooth", in reference to the shape) was a herbivorous genus of sauropod dinosaur, based on a tooth from the late Bathonian-age Middle Jurassic Forest Marble Formation of Wiltshire, England. Historically, it is very obscure and usually referred to Cetiosaurus , but recent analyses suggest that it is a distinct genus, and possibly related to Turiasaurus . Cardiodon was the first sauropod genus named. [1]
Richard Owen named the genus for a now-lost tooth, part of the collection of naturalist Joseph Chaning Pearce, found near Bradford-on-Avon, but did not assign it a specific name at the time. The generic name is derived from Greek καρδία, kardia, "heart", and ὀδών, odon, "tooth", in reference to its heart-shaped profile. [2] A few years later, in 1844, he added the specific name rugulosus, meaning "wrinkled" in Latin. [3] Cardiodon was the first sauropod given a formal name to, though Owen was at the time completely unaware of the sauropod nature of the find.
Within a few decades, he and others were viewing Cardiodon as a possible synonym of his most well-known sauropod genus, Cetiosaurus . [4] [5] Richard Lydekker formalized this view in a roundabout way in 1890, by assigning Cetiosaurus oxoniensis to Cardiodon on the basis of teeth from Oxfordshire associated with a skeleton of C. oxoniensis. [6] He also added a second tooth (BMNH R1527) from the Great Oolite near Cirencester, Gloucestershire. [6] More typically, Cardiodon has been assigned to Cetiosaurus, sometimes as a separate species Cetiosaurus rugulosus, [7] in spite of its priority.
In 2003, Paul Upchurch and John Martin, reviewing Cetiosaurus, found that there is little evidence to assign the C. oxoniensis teeth to the skeleton, and the "C. oxoniensis" teeth differ from the Cardiodon teeth (Cardiodon teeth are convex facing the tongue); therefore, they supported Cardiodon being retained as its own genus. [8] Upchurch et al. (2004) repeated this assessment, and found that though the teeth have no known autapomorphies, they are those of a eusauropod. [9] More recently, Royo-Torres et al. (2006), in their description of Turiasaurus , pointed out Cardiodon as a possible relative to their new, giant sauropod, placing it in the Turiasauria. [10] Earlier, Cardiodon had been usually assigned to the Cetiosauridae or a Cardiodontidae of its own.
The original tooth shows, as far as can be deduced from the surviving illustrations, the rare combination of being spatulate and having a convex inner side, though the convexity is slight. Its crown is short and wide, slightly curving to the inside. The outer side is strongly convexly curved from the front to the rear. On this side a shallow groove is present, running parallel to the rear edge. The crown tapers towards its tip. The edges have no denticles. The enamel shows the little wrinkles to which the specific name refers. [8]
As a sauropod, Cardiodon would have been a large, quadrupedal herbivore, [9] but because of the scanty remains, much more cannot be said.
Pelorosaurus is a genus of titanosauriform sauropod dinosaur. Remains referred to Pelorosaurus date from the Early Cretaceous period, about 140-125 million years ago, and have been found in England and Portugal. Thomas Holtz estimated its length at 24 meters.
Aepisaurus was a genus of sauropod dinosaur from the Albian-age Lower Cretaceous Grès vert of Département du Vaucluse, France, around 100.5 million years ago. It is an obscure genus from an unknown family, represented by a single humerus, now partly lost. Despite its lack of popularity, or perhaps because of it, it has been misspelled several ways in the scientific literature, with multiple dates given to the year of description as well.
Cetiosaurus meaning 'whale lizard', from the Greek keteios/κήτειος meaning 'sea monster' and sauros/σαυρος meaning 'lizard', is a genus of herbivorous sauropod dinosaur from the Middle Jurassic Period, living about 168 million years ago in what is now Britain and probably France.
Cetiosauriscus is a genus of sauropod dinosaur that lived between 166 and 164 million years ago during the Callovian in what is now England. A herbivore, Cetiosauriscus had – by sauropod standards – a moderately long tail, and longer forelimbs, making them as long as its hindlimbs. It has been estimated as about 15 m (49 ft) long and between 4 and 10 t in weight.
Haplocanthosaurus is a genus of intermediate sauropod dinosaur. Two species, H. delfsi and H. priscus, are known from incomplete fossil skeletons. It lived during the late Jurassic period, 155 to 152 million years ago. The type species is H. priscus, and the referred species H. delfsi was discovered by a young college student named Edwin Delfs in Colorado, United States. Haplocanthosaurus specimens have been found in the very lowest layer of the Morrison Formation, along with Hesperosaurus mjosi, Brontosaurus yahnahpin, and Allosaurus jimmadseni.
Dinodocus is a genus of sauropod dinosaur, named by Richard Owen in 1884. The name is now usually considered a nomen dubium. The only species, D. mackesoni, a name given to some fossil bones from the Lower Greensand Group of Hythe, Kent, England, were formerly placed in the genus Pelorosaurus, but a review by Upchurch et al. (2004) concluded that Dinodocus is a nomen dubium.
Rebbachisauridae is a family of sauropod dinosaurs known from fragmentary fossil remains from the Cretaceous of South America, Africa, North America, Europe and possibly Central Asia.
Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.
Eucamerotus was a genus of sauropod dinosaur from the Barremian-age Lower Cretaceous Wessex Formation (Wealden) of the Isle of Wight, England.
Neosodon was a genus of sauropod dinosaur from the Late Tithonian-age Upper Jurassic Sables et Gres a Trigonia gibbosa of Pas-de-Calais department, France. It has never been formally given a species name, but is often seen as N. praecursor, which actually comes from a different animal. Often in the past, it had been assigned to the wastebasket taxon Pelorosaurus, but restudy has suggested that it could be related to Turiasaurus, a roughly contemporaneous giant Spanish sauropod. It is only known from six teeth.
Oplosaurus was a genus of sauropod dinosaur from the Barremian-age Lower Cretaceous Wessex Formation of the Isle of Wight, England. It is known from a single tooth usually referred to the contemporaneous "wastebasket taxon" Pelorosaurus, although there is no solid evidence for this.
Ornithopsis is a genus of sauropod dinosaur, from the Early Cretaceous of England and possibly Germany. The type species, which is the only species seen as valid today, is O. hulkei, which is only known from fragmentary remains.
Lapparentosaurus is a genus of sauropod dinosaur from the Middle Jurassic. Its fossils were found in Madagascar. The type species is L. madagascariensis.
Turiasauria is an unranked clade of basal sauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa.
Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thick jaw. Based on cervical vertebrae and cervical rib biomechanics, camarasaurids most likely moved their necks in a vertical, rather than horizontal, sweeping motion, in contrast to most diplodocids.
Suchosaurus is a spinosaurid dinosaur from Cretaceous England and Portugal, originally believed to be a genus of crocodile. The type material, consisting of teeth, was used by British palaeontologist Richard Owen to name the species S. cultridens in 1841. Later in 1897, French palaeontologist Henri-Émile Sauvage named a second species, S. girardi, based on two fragments from the mandible and one tooth discovered in Portugal. Suchosaurus is possibly a senior synonym of the contemporary spinosaurid Baryonyx, but is usually considered a dubious name due to the paucity of its remains, and is considered an indeterminate baryonychine. In the Wadhurst Clay Formation of what is now southern England, Suchosaurus lived alongside other dinosaurs, as well as plesiosaurs, mammals, and crocodyliforms.
Neosauropoda is a clade within Dinosauria, coined in 1986 by Argentine paleontologist José Bonaparte and currently described as Saltasaurus loricatus, Diplodocus longus, and all animals directly descended from their most recent common ancestor. The group is composed of two subgroups: Diplodocoidea and Macronaria. Arising in the early Jurassic and persisting until the Cretaceous–Paleogene extinction event, Neosauropoda contains the majority of sauropod genera, including genera such as Apatosaurus, Brachiosaurus, and Diplodocus. It also includes giants such as Argentinosaurus, Patagotitan and Sauroposeidon, and its members remain the largest land animals ever to have lived.
Duriatitan is a genus of titanosauriform sauropod dinosaur that lived in the Late Jurassic in what is now England. The holotype specimen of Duriatitan, BMNH 44635, is a partial left upper arm bone which was found by R.I. Smith near Sandsfoot, Weymouth in the lower Kimmeridge Clay from Dorset. The type species, D. humerocristatus, was described in 1874 by John Hulke as a species of Cetiosaurus and was noted as being similar to that of Gigantosaurus. The specific name refers to the deltopectoral crest, crista, on the upper arm bone, humerus. The specimen was assigned to its own genus by Paul M. Barrett, Roger B.J. Benson and Paul Upchurch in 2010. The generic name is derived from the Latin name for Dorset, Duria, and Greek Titan. Thomas Holtz estimated its length at 25 meters (82 ft).
Haestasaurus is a genus of herbivorous sauropod dinosaur, belonging to the Macronaria, that during the Early Cretaceous lived in the area of present-day England. The only species is Haestasaurus becklesii.
Mierasaurus is an extinct genus of sauropod dinosaur from the Early Cretaceous of Utah, United States. The taxon was first described and named in 2017 by Rafael Royo-Torres and colleagues, from a mostly complete skeleton including a disarticulated partial skull and mandible, teeth, multiple vertebrae from along the length of the body, both scapulae, radius and ulna bones, a left manus, a complete pelvis, both femora and the entire left hindlimb. Additionally, they referred a lower jaw and femur from juvenile individuals, which were found nearby, to the genus. Collectively, Mierasaurus is among the most completely known North American sauropods. The genus name honours Bernardo de Miera y Pacheco, the first European scientist to enter what is now Utah. The type species for Mierasaurus is Mierasaurus bobyoungi, named after Robert Glen Young, a paleontologist who researched the Early Cretaceous of Utah.