Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan—estimated at 37 m (121 ft) long with a weight of 69 tonnes —and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
The Brachiosauridae are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the leaves of tall trees that other sauropods would have been unable to reach. In addition, they possessed thick spoon-shaped teeth which helped them to consume tough plants more efficiently than other sauropods. They have also been characterized by a few unique traits or synapomorphies; dorsal vertebrae with 'rod-like' transverse processes and an ischium with an abbreviated pubic peduncle.
Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species A. maximus was transferred over to the new genus Arrudatitan in 2021.
Euhelopus is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian stages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Like sauropods such as brachiosaurs and titanosaurs, Euhelopus had longer forelegs than hind legs. This discovery was paleontologically significant because it represented the first dinosaur scientifically investigated from China: seen in 1913, rediscovered in 1922, and excavated in 1923 and studied by T'an during the same year. Unlike most sauropod specimens, it has a relatively complete skull.
Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.
Cedarosaurus was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period (Valanginian). It was a sauropod which lived in what is now Utah. The fossils were discovered in 1996 in eastern Utah within the rocks of the Yellow Cat Member of the Cedar Mountain Formation. It was officially named and described by Tidwell, Carpenter and Brooks in 1999.
Huabeisaurus was a genus of dinosaur from the Late Cretaceous. It was a sauropod which lived in what is present-day northern China. The type species, Huabeisaurus allocotus, was first described by Pang Qiqing and Cheng Zhengwu in 2000. Huabeisaurus is known from numerous remains found in the 1990s, which include teeth, partial limbs and vertebrae. Due to its relative completeness, Huabeisaurus represents a significant taxon for understanding sauropod evolution in Asia. Huabeisaurus comes from Kangdailiang and Houyu, Zhaojiagou Town, Tianzhen County, Shanxi province, China. The holotype was found in the unnamed upper member of the Huiquanpu Formation, which is Late Cretaceous (?Cenomanian–?Campanian) in age based on ostracods, charophytes, and fission-track dating.
Sonorasaurus is a genus of brachiosaurid dinosaur from the Early to Late Cretaceous. It was a herbivorous sauropod whose fossils have been found in southern Arizona in the United States. Its name, which means "Sonora lizard", comes from the Sonora River that flows in the Sonoran Desert where its fossils were first found. The type species is S. thompsoni, described by Ratkevich in 1998.
Venenosaurus is a genus of sauropod dinosaur that lived in what is now Utah during the Early Cretaceous. Its type and only species is Venenosaurus dicrocei. Fossils of Venenosaurus were first discovered in 1998, by Denver Museum of Natural History volunteer Anthony DiCroce, and described as a new genus and species in 2001 by Virginia Tidwell and colleagues, who named the species for DiCroce. Venenosaurus was a relatively small sauropod, and was similar to Cedarosaurus, another sauropod from the Early Cretaceous of Utah.
Losillasaurus is a genus of sauropod dinosaur from the Late Jurassic and possibly Early Cretaceous (Kimmeridgian-?Berriasian) in the southeast of Spain. The type species of the turiasaurian Losillasaurus giganteus was discovered in the Villar del Arzobispo Formation in Valencia and formally described by Casanovas, Santafé and Sanz in 2001. The holotype material is from a subadult and includes part of a skull; complete cervical, dorsal, sacral, and caudal vertebrae as well as several fragments; skeletal elements from the limbs including a humerus, ulna, radius, and metacarpal; sternal plates; and from the pelvis: the ilium, ischium, and pubis. The genus is characterized by the dimension and shape of the neural spine of the proximal caudal vertebrae. The humerus is 143 centimetres (56 in) long, which despite being from a subadult specimen is within 20% of the size of Paralititan. The size estimation proposed by Francisco Gascó in his master thesis is 15–18 m (49–59 ft) and 12-15 tons.
Europasaurus is a basal macronarian sauropod, a form of quadrupedal herbivorous dinosaur. It lived during the Late Jurassic of northern Germany, and has been identified as an example of insular dwarfism resulting from the isolation of a sauropod population on an island within the Lower Saxony basin.
Amargatitanis is a genus of dicraeosaurid sauropod dinosaur from the Barremian-age La Amarga Formation of Neuquén, Argentina. It is known from a single, incomplete postcranial skeleton consisting of a partial hindlimb, ischium, and two vertebrae. These remains were unearthed by Argentine paleontologist José Bonaparte in 1983 during an expedition by the Museo Argentino de Ciencias Naturales and later described as a new genus and species, Amargatitanis macni by Sebastián Apesteguía. The genus name comes from the words Amarga, where the fossils were collected, and titanis meaning "titan". Its species name is in reference to the MACN, where the remains are stored.
Tastavinsaurus is a genus of sauropod dinosaur belonging to the Titanosauriformes. It is based on a partial skeleton from the Early Cretaceous Xert Formation of Spain. The type species is Tastavinsaurus sanzi, named in honor of the Rio Tastavins in Spain and Spanish paleontologist José Luis Sanz. Tastavinsaurus sanzi is known from two specimens, one from the Xert Formation and one from the Forcall Formation. Tastavinsaurus was originally described as a somphospondylan, but it may be a brachiosaurid. The clade Laurasiformes was coined for Tastavinsaurus and its close relatives, which may include Cedarosaurus, Soriatitan, and Venenosaurus.
Brachiosaurus is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154 to 150 million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in the Colorado River valley in western Colorado, United States. Riggs named the dinosaur Brachiosaurus altithorax; the generic name is Greek for "arm lizard", in reference to its proportionately long arms, and the specific name means "deep chest". Brachiosaurus is estimated to have been between 18 and 22 meters long; body mass estimates of the subadult holotype specimen range from 28.3 to 46.9 metric tons. It had a disproportionately long neck, small skull, and large overall size, all of which are typical for sauropods. Atypically, Brachiosaurus had longer forelimbs than hindlimbs, which resulted in a steeply inclined trunk, and a proportionally shorter tail.
Normanniasaurus is an extinct genus of basal titanosaur sauropod known from the Early Cretaceous Poudingue Ferrugineux of Seine-Maritime, northwestern France.
Padillasaurus is an extinct genus of titanosauriform sauropod known from the Early Cretaceous Paja Formation in Colombia. It contains a single species, Padillasaurus leivaensis, known only from a single partial axial skeleton. Initially described as a brachiosaurid, it was considered to be the first South American brachiosaurid ever discovered and named. Before its discovery, the only known brachiosaurid material on the continent was very fragmentary and from the Jurassic period. However, a more recent study finds it to be a basal somphospondylan.
Savannasaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Winton Formation of Queensland, Australia. It contains one species, Savannasaurus elliottorum, named in 2016 by Stephen Poropat and colleagues. The holotype and only known specimen, originally nicknamed "Wade", is the most complete specimen of an Australian sauropod, and is held at the Australian Age of Dinosaurs museum. Dinosaurs known from contemporary rocks include its close relative Diamantinasaurus and the theropod Australovenator; associated teeth suggest that Australovenator may have fed on the holotype specimen.
Magnamanus is an extinct genus of herbivorous iguanodontian dinosaur that lived during the Early Cretaceous in what is now Spain in the Golmayo Formation. It contains a single species, Magnamanus soriaensis.
Vouivria is a genus of herbivorous sauropod dinosaur, belonging to the Brachiosauridae, that lived in the area of present France during the Late Jurassic. The type species is Vouivria damparisensis.
Garumbatitan is an extinct genus of somphospondylan sauropod dinosaur from the Cretaceous Arcillas de Morella Formation of Spain. The genus contains a single species, G. morellensis, known from multiple partial skeletons.
