Kholumolumo Temporal range: Late Triassic, | |
---|---|
Pes of K. ellenbergerorum (specimen NMQR1705) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Massopoda |
Genus: | † Kholumolumo Peyre de Fabrègues & Allain, 2020 |
Type species | |
†Kholumolumo ellenbergerorum Peyre de Fabrègues & Allain, 2020 |
Kholumolumo (referring to a type of dragon the local Basuto associate with dinosaurs), formerly "Kholumolumosaurus" or "Thotobolosaurus", is an extinct genus of massopodan sauropodomorph dinosaur, which was closely related to Sarahsaurus , [1] from the lower Elliot Formation of Maphutseng, Lesotho. The type species, Kholumolumo ellenbergerorum was formally described in 2020. [1]
In 1930, Samuel Motsoane, principal of the Paris Evangelical Mission School at Bethesda in Lesotho, found dispersed dinosaur bones. In 1955, he told this to the protestant missionary Paul Ellenberger. In September 1955, P. Ellenberger and his brother François Ellenberger uncovered a bonebed in the immediate vicinity of a refuse pile a few meters from native huts in the Village of Maphutseng, western Lesotho, in a layer of the Elliot Formation. The location was locally called the Thotobolo ea ‘Ma-Beata, the "trash heap of Beata's mother". In November 1955, they were reinforced by the South-African paleontologists Alfred Walter Crompton and Rosalie F. "Griff" Ewer. [1] In 1955, the discovery was reported in the scientific literature. [2] The excavations were continued from February 1956 onwards and at the end of the second field season, the number of pieces had increased to 683, collected from a surface of thirty-five square metres. The Ellenberger brothers briefly described the finds in 1956. [3] In 1957, the fossils were transported to the South African Museum of Cape Town. Delays in their preparation caused a rift between Crompton and the Ellenbergers. [1]
In 1959, P. Ellenberger, F. Ellenberger and the latter's wife Hélène, continued the digs in cooperation with a French team from the Paris Muséum national d'histoire naturelle including Léonard Ginsburg and Jean Fabre. Over two hundred additional pieces were collected. [1] In 1960, eight dinosaur trackways were reported, discovered over a surface of seventy square metres. [4] Three individual tracks were transported to the University of Montpellier but are presently lost. [1] In 1963, additional excavations took place by Paul Ellenberger, Ginsburg, Fabre and Christiane Mendrez. [1] In 1966, François Ellenberger and Ginsburg for the first time described the bones in detail and referred them to Euskelosaurus browni . [5]
In September 1970, the last excavations occurred by Paul Ellenberger, Ginsburg, Fabre and Bernard Battail. As in 1959, the fossils were sent to Paris, bringing the total there to about four hundred. [1] Concluding that it represented a taxon new to science, Paul Ellenberger at first referred to it as "Thotobolosaurus", which means "trash heap lizard", in reference to where the holotype was discovered. Ellenberger in a redescription of the material suggested the species name "Thotobolosaurus mabeatae" in 1970. [6] This name was invalid, however. [1]
In 1996, the species was briefly referred to as "Kholumolumosaurus ellenbergerorum" in François-Xavier Gauffre's unpublished dissertation. [7] [8] This name too, was invalid. [1]
In 2020, the species Kholumolumo ellenbergerorum, a shortened form of one of the original names, was formally named by Claire Peyre de Fabrègues and Ronan Allain. The generic name is the kholumolumo or xodumodumo, a gigantic reptile, sometimes described as a dragon, lizard or crocodile, from the mythology of the Sotho. The Basotho use this word to refer to dinosaurs. The specific name honours the Ellenberger family. [1]
The holotype, MNHN.F.LES381m, was found in a layer of the Lower Elliot Formation dating from the Norian. It consists of a complete right tibia or shinbone. Several other bones in the Paris material were designated as paratypes. All the additional sauropodomorph material from Maphutseng, whether in France or South-Africa, was referred to the species. Only disarticulated bones were found, including little skull material. They represent at least five, probably about ten, individuals. [1]
Kholumolumo was a large sauropodomorph, estimated to have reached upwards of 9 metres (30 ft) long. [1] This would have made it one of the largest known animals of the Norian. [1] Its weight has been estimated from its thighbone circumference. If it was bipedal, which the describing authors thought likely, a weight is indicated of 1754 kilogrammes. A quadrupedal animal could have weighed 3963 kilogrammes. [1]
Since no complete skeletons of Kholumolumo have been discovered, much of what is known about its physical appearance and diet has to be inferred from its close relatives. Most likely it would be a herbivore that looked similar to dinosaurs such as Sarahsaurus.
The holotype shows a unique combination of traits that in themselves are not unique. The shinbone is very short and robust, its circumference measuring 53% of its length: in all other known non-sauropod Sauropodomorpha this ratio is lower than 0.49, with the exception of Antetonitrus and Blikanasaurus . The shinbone shaft tapers to below, both seen from the inside as viewed from the outside. The shinbone shaft has a straight front and rear edge, different from Antetonitrus. [1]
Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.
Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. An omnivore from South Africa, it had a large body and sturdy limbs, suggesting it moved quadrupedally. Its limb bones were massive and heavy like the limb bones of true sauropods.
Fabrosaurus is a dubious extinct genus of ornithischian dinosaur that lived during the Early Jurassic during the Hettangian to Sinemurian stages 199 - 189 mya. It was originally placed within the now obsolete family Fabrosauridae.
Isanosaurus is an extinct genus of sauropod dinosaur from Thailand. It was originally dated to approximately 210 million years ago during the Late Triassic, which would make it one of the oldest known sauropods. Its age was later considered uncertain, and may be Early Jurassic or even as young as Late Jurassic. The only species is Isanosaurus attavipachi. Though important for the understanding of sauropod origin and early evolution, Isanosaurus is poorly known. Exact relationships to other early sauropods remain unresolved.
Antetonitrus is a genus of sauropodiform dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is Antetonitrus ingenipes. Sometimes considered a basal sauropod, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support.
Basutodon was a genus of suchian archosaur from the late Carnian-early Norian-age Upper Triassic Lower Elliot Formation of Lesotho. The type species is B. ferox.
Euskelosaurus is a sauropodomorph dinosaur from the Late Triassic of South Africa and Lesotho. Fossils have only been recovered from the lower Elliot Formation in South Africa and Lesotho, and in one locality in Zimbabwe.
Gryponyx is an extinct genus of massopod sauropodomorph known from southern Free State, central South Africa.
Lusitanosaurus is a genus of reptile from the Sinemurian stage of Early Jurassic of Portugal, maybe from the Coimbra Formation. It was considered the second example of the Dinosaurian group Thyreophora from the Sinemurian of Europe and it the oldest known dinosaur from the Iberian Peninsula, but this affinity has been contested. It is based on a large left maxilla with teeth that was lost in the fire at Museu Nacional de História Natural e da Ciência, Lisbon, in 1978.
Inosaurus is the name given to a dubious genus of theropod dinosaur from the Early Cretaceous-aged "Continental intercalaire" and Early Cretaceous (Albian)-aged Echkar Formation of Niger and possibly from the Late Cretaceous (Cenomanian)-aged Baharija Formation of Egypt The type, and only species, is Inosaurus tedreftensis.
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town.
The Assaouas Formation, also referred to as the Assaouas Sandstone is a Late Jurassic geologic formation in Niger. Fossil sauropod tracks have been reported from the formation.
Ignavusaurus is a genus of basal sauropodomorph dinosaur that lived during the Early Jurassic in what is now Lesotho. Its fossils were found in the Upper Elliot Formation which is probably Hettangian in age. It was described on the basis of a partial, well preserved articulated skeleton. The type species, I. rachelis, was described in 2010 by Spanish palaeontologist F. Knoll.
The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group rocks are considered to range between Lower Triassic (Olenekian) to Lower Jurassic (Pliensbachian) in age. These estimates are based on means of geological dating including stratigraphic position, lithostratigraphic and biostratigraphic correlations, and palynological analyses.
Pegomastax is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic of South Africa. The only known specimen was discovered in a 1966–1967 expedition in Transkei District of Cape Province, but was not described until 2012 when Paul Sereno named it as the new taxon Pegomastax africana. The genus name is derived from the Greek for "strong jaw", and the species name describes the provenance of Africa; it was originally spelled africanus, was corrected to africana to align with the gender of the genus name.
Pulanesaura is an extinct genus of basal sauropodiform known from the Early Jurassic Upper Elliot Formation of the Free State, South Africa. It contains a single species, Pulanesaura eocollum, known from partial remains of at least two subadult to adult individuals.
Meroktenos is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period of what is now Lesotho.
Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.
Ngwevu is a genus of massospondylid sauropodomorph dinosaur from the Lower Jurassic of South Africa. The genus contains one species, Ngwevu intloko.