Menucocelsior Temporal range: Late Cretaceous, | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Clade: | † Eutitanosauria |
Genus: | † Menucocelsior Rolando et al. 2022 |
Type species | |
†Menucocelsior arriagadai Rolando et al., 2022 |
Menucocelsior (meaning "the taller one from Salitral Ojo del Agua") is a genus of medium-sized titanosaurian sauropod dinosaur from the Late Cretaceous Allen Formation of Argentina. The type and only species is Menucocelsior arriagadai.
The Salitral Ojo del Agua area of Río Negro, Argentina was home to an entire sauropod fauna during the late Cretaceous period. Fossil material belonging to saltasaurines, aeolosaurines, and other titanosaurs have been discovered. In 2022, Rolando et al. designated one specimen as the type species for a new titanosaurian taxon, Menucocelsior arriagadai. The generic name combines the Mapudungun menuco, meaning "watering hole", a translation of Ojo del Agua, with the Latin celsior, meaning "higher" or "major." The specific name honours "Beto" Arriagada, the owner of the ranch where the discoveries were made. [1]
The holotype, MPCN-PV-798, consists of seventeen caudal vertebrae and several appendicular bones: a right humerus, a left fibula and some metapodial. The structure and form of these bones suggests it was a member of the Eutitanosauria, but not a member of the Colossosauria, Saltasaurinae, or Aeolosaurini. Fossil material suggests that Menucocelsior was about 8 metres (26 ft) long. [1]
Menucocelsior lived in the Allen Formation alongside a number of other titanosaurs, including the small saltasaurine Rocasaurus and indeterminate saltasaurines and aeolosaurins, known from both skeletal remains and differing morphotypes of osteoderms. The describers note that such an assemblage of titanosaurs is not known anywhere else in the world; the titanosaurs' differing body plans probably allowed them to occupy differing ecological niches and thus limit competition. [1]
Saltasaurus is a genus of saltasaurid dinosaur of the Late Cretaceous period of Argentina. Small among sauropods, though still heavy by the standards of modern creatures, Saltasaurus was characterized by a short neck and stubby limbs. It was the first genus of sauropod known to possess armour of bony plates embedded in its skin. Such small bony plates, called osteoderms, have since been found on other titanosaurians.
Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan—estimated at 37 m (121 ft) long with a weight of 69 tonnes —and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
Antarctosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. The type species, Antarctosaurus wichmannianus, and a second species, Antarctosaurus giganteus, were described by prolific German paleontologist Friedrich von Huene in 1929. Three additional species of Antarctosaurus have been named since then but later studies have considered them dubious or unlikely to pertain to the genus.
Alamosaurus is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to Argentinosaurus and Puertasaurus, which would make it the absolute largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age. Specimens of a juvenile Alamosaurus sanjuanensis have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Alamosaurus is the only known sauropod to have inhabited North America after their nearly 30-million year absence from the North American fossil record and probably represents an immigrant from South America.
Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species A. maximus was transferred over to the new genus Arrudatitan in 2021.
Bonitasaura is a genus of titanosaurian dinosaur hailing from uppermost layers of the Late Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén Group of the eastern Neuquén Basin, located in Río Negro Province, Northwestern Patagonia, Argentina. The remains, consisting of a partial sub-adult skeleton jumbled in a small area of fluvial sandstone, including a lower jaw with teeth, a partial vertebrae series, and limb bones, were described by Sebastian Apesteguía in 2004.
Clasmodosaurus is a genus of titanosaurian sauropod dinosaur from the Bajo Barreal Formation. It lived during the Late Cretaceous in what is now Argentina. It is known only from three fossilized teeth, and is therefore a tooth taxon. It is a nomen dubium.
Isisaurus is a genus of titanosaurian dinosaur from the Late Cretaceous Lameta Formation of India and Pab Formation of Pakistan. The genus contains a single species, Isisaurus colberti.
Neuquensaurus is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived.
Rocasaurus is a genus of titanosaurian sauropod that lived in South America. Rocasaurus was discovered in Argentina in 2000, within the Allen Formation which is dated to be middle Campanian to early Maastrichtian in age. This genus grew up to 8 metres (26 ft) long, making it one of the smaller sauropods. It seems to be closely related to saltasaurid dinosaurs, like Saltasaurus and Neuquensaurus.
Pellegrinisaurus is a genus of titanosaurian sauropod dinosaur that lived in South America during the Late Cretaceous period. The holotype was found in the Allen Formation, Argentina.
The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous (middle Campanian to early Maastrichtian. Dinosaur remains are among the fossils that have been recovered from the formation. Indeterminate chelid remains and other vertebrates have also been discovered in this formation.
Bonapartenykus is a monospecific genus of alvarezsauroid dinosaur from Argentina that lived during the Late Cretaceous (Campanian-Maastrichtian) in what is now the upper Allen Formation of the Río Negro Province. The type and only species, Bonapartenykus ultimus, is known from a nearly articulated but partial skeleton that was found in close association to two incomplete eggs and several clusters of eggshells belonging to the oogenus Arriagadoolithus. Bonapartenykus was named in 2012 by Federico L. Agnolin, Jaime E. Powell, Fernando E. Novas and Martin Kundrát. Bonapartenykus has an estimated length of 2.5 m (8.2 ft) and weight of 72 kg (159 lb), making it the largest member of the clade Alvarezsauroidea.
Eutitanosauria is a clade of titanosaurs, encompassing the more derived members of the group and characterized by the absence of the hyposphene-hypantrum articulation and possibly the presence of osteoderms. The group was first named by Sanz and colleagues in 1999, who used it to unite the group of Argyrosaurus, Lirainosaurus, Saltasaurus and the Peiropolis titanosaur. However, this definition was not used as it made the group equivalent to Saltasauridae, so Saldago redefined it in 2003 to be all titanosaurs closer to Saltasaurus than Epachthosaurus. This definition created Eutitanosauria as the sister group to Epachthosaurinae, but was problematic due to the variable nature of Epachthosaurus. Eutitanosauria was often broadly similar to Lithostrotia, and has often been unused or unlabelled on phylogenies. Sometimes Epachthosaurus would be more primitive than Malawisaurus, making Eutitanosauria more encompassing than Lithostrotia, or Epachthosaurus could nest close to Colossosauria and limit Eutitanosauria to a smaller group of saltasauroids. Because of the flexible nature of Epachthosaurus in basal titanosaur phylogeny, Carballido and colleagues redefined the group in 2022 to include the smallest clade of both Patagotitan, a colossosaur, and Saltasaurus, creating a node-stem clade with Colossosauria and Saltasauroidea, presenting the informal cladogram of stable titanosaur clades below.
Niebla is a genus of abelisaurid theropod dinosaur from the Late Cretaceous Period (Campanian-Maastrichtian) of Río Negro province, Argentina. The genus contains a single species, Niebla antiqua, and is known from a partial, non-articulated skeleton. The holotype, found in the Allen Formation, represents an adult individual.
Abditosaurus is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia, Spain. The type and only species is Abditosaurus kuehnei. Phylogenetic analyses recover it within a clade of South American and African saltasaurines, distinct from other insular dwarf sauropods from the European archipelago. Abditosaurus inhabited the Ibero-Armorican Island, a prehistoric island made up of what is now Spain, Portugal, and southern France, and would have been the largest titanosaur species in its environment.
Ibirania is a genus of dwarf saltasaurine titanosaur dinosaur from the Late Cretaceous São José do Rio Preto Formation of Southeast Brazil. The type species is Ibirania parva. It is one of the smallest sauropods known to date, comparable in size to the titanosaur Magyarosaurus.
Patagopelta is an extinct genus of ankylosaurian dinosaur from the Late Cretaceous Allen Formation of Argentina. The genus contains a single species, P. cristata, known from a partial skeleton. While originally described as a nodosaurine, later discoveries provided support for parankylosaurian affinities for the taxon. Patagopelta is a very small ankylosaur, comparable in size to the dwarf nodosaurid Struthiosaurus, about 2 m (6.6 ft) long.
Sidersaura is an extinct genus of rebbachisaurid sauropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, S. marae, known from the remains of four individuals. Sidersaura represents one of the largest known rebbachisaurids.
Udelartitan is an extinct genus of saltasauroid titanosaurian sauropod dinosaur from the Late Cretaceous Guichón Formation of Uruguay. The genus contains a single species, U. celeste, known from fragmentary remains of at least two individuals.