The prefrontal bone is a bone separating the lacrimal and frontal bones in many tetrapod skulls. It first evolved in the sarcopterygian clade Rhipidistia, which includes lungfish and the Tetrapodomorpha. [1] The prefrontal is found in most modern and extinct lungfish, amphibians and reptiles. The prefrontal is lost in early mammaliaforms and so is not present in modern mammals either. [2]
The prefrontal bone is a very small bone near the top of the skull, which is lost in many groups of coelurosaurian theropod dinosaurs and is completely absent in their modern descendants, the birds. Conversely, a well developed prefrontal is considered to be a primitive feature in dinosaurs. The prefrontal makes contact with several other bones in the skull. The anterior part of the bone articulates with the nasal bone and the lacrimal bone. The posterior part of the bone articulates with the frontal bone and more rarely the palpebral bone. The prefrontal bone sometimes forms part of the upper margin of the orbit. This bone is part of the skull roof, which is a set of bones that cover the brain, eyes and nostrils.
Tetrapods are four-limbed animals constituting the superclass Tetrapoda. It includes extant and extinct amphibians, reptiles, and synapsids. Tetrapods evolved from a group of animals known as the Tetrapodomorpha which, in turn, evolved from ancient sarcopterygian fish around 390 million years ago in the middle Devonian period; their forms were transitional between lobe-finned fishes and the four-limbed tetrapods. The first crown-tetrapods appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors of the tetrapods and the process by which they colonized Earth's land after emerging from water remains unclear. The change from a body plan for breathing and navigating in water to a body plan enabling the animal to move on land is one of the most profound evolutionary changes known. The first tetrapods (stem) or "fishapods" were primarily aquatic. Modern amphibians, which evolved from earlier groups, are generally semiaquatic; the first stage of their lives is as fish-like tadpoles, and later stages are partly terrestrial and partly aquatic. However, most tetrapod species today are amniotes, most of which are terrestrial tetrapods whose branch evolved from earlier tetrapods about 340 million years ago. The key innovation in amniotes over amphibians is the amnion, which enables the eggs to retain their aqueous contents on land, rather than needing to stay in water.
The skull is a bone structure that forms the head in vertebrates. It supports the structures of the face and provides a protective cavity for the brain. The skull is composed of two parts: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton.
Lungfish are freshwater rhipidistian vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods.
Sarcopterygii — sometimes considered synonymous with Crossopterygii — is a taxon of the bony fishes whose members are known as lobe-finned fishes. The group Tetrapoda, a superclass including amphibians, reptiles, and mammals, evolved from certain sarcopterygians; under a cladistic view, tetrapods are themselves considered a group within Sarcopterygii.
The frontal bone is a bone in the human skull. The bone consists of two portions. These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, part of the bony orbital cavity holding the eye, and part of the bony part of the nose respectively. The name comes from the Latin word frons.
The parietal bones are two bones in the skull which, when joined at a fibrous joint, form the sides and roof of the cranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, and four angles. It is named from the Latin paries (-ietis), wall.
Anders Erik Vilhelm Jarvik was a Swedish paleontologist who worked extensively on the sarcopterygian fish Eusthenopteron. In a career that spanned some 60 years, Jarvik produced some of the most detailed anatomical work on this fish, making it arguably the best known fossil vertebrate.
The lacrimal bone is a small and fragile bone of the facial skeleton; it is roughly the size of the little fingernail. It is situated at the front part of the medial wall of the orbit. It has two surfaces and four borders. Several bony landmarks of the lacrimal bone function in the process of lacrimation or crying. Specifically, the lacrimal bone helps form the nasolacrimal canal necessary for tear translocation. A depression on the anterior inferior portion of the bone, the lacrimal fossa, houses the membranous lacrimal sac. Tears or lacrimal fluid, from the lacrimal glands, collect in this sac during excessive lacrimation. The fluid then flows through the nasolacrimal duct and into the nasopharynx. This drainage results in what is commonly referred to a runny nose during excessive crying or tear production. Injury or fracture of the lacrimal bone can result in posttraumatic obstruction of the lacrimal pathways.
The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians.
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species.
Panderichthys is a genus of extinct sarcopterygian from the late Devonian period, about 380 Mya. Panderichthys, which was recovered from Frasnian deposits in Latvia, is represented by two species. P. stolbovi is known only from some snout fragments and an incomplete lower jaw. P. rhombolepis is known from several more complete specimens. Although it probably belongs to a sister group of the earliest tetrapods, Panderichthys exhibits a range of features transitional between tristichopterid lobe-fin fishes and early tetrapods. It is named after the German-Baltic paleontologist Christian Heinrich Pander. Possible tetrapod tracks dating back to before the appearance of Panderichthys in the fossil record were reported in 2010, which suggests that Panderichthys is not a direct ancestor of tetrapods, but nonetheless shows the traits that evolved during the fish-tetrapod evolution
Dipnorhynchus is an extinct genus of lungfish from the middle Devonian period of Australia and Europe.
Jesairosaurus is an extinct genus of early archosauromorph reptile known from the Illizi Province of Algeria. It is known from a single species, Jesairosaurus lehmani. Although a potential relative of the long-necked tanystropheids, this lightly-built reptile could instead be characterized by its relatively short neck as well as various skull features.
Sauroniops is a genus of carnivorous basal carcharodontosaurid theropod dinosaur known from the Late Cretaceous of Morocco.
This glossary explains technical terms commonly employed in the description of dinosaur body fossils. Besides dinosaur-specific terms, it covers terms with wider usage, when these are of central importance in the study of dinosaurs or when their discussion in the context of dinosaurs is beneficial. The glossary does not cover ichnological and bone histological terms, nor does it cover measurements.
Innovations conventionally associated with terrestrially first appeared in aquatic elpistostegalians such as Panderichthys rhombolepis, Elpistostege watsoni, and Tiktaalik roseae. Phylogenetic analyses distribute the features that developed along the tetrapod stem and display a stepwise process of character acquisition, rather than abrupt. The complete transition occurred over a period of 30 million years beginning with the tetrapodomorph diversification in the Middle Devonian.
Boreopricea is an extinct genus of archosauromorph reptile from the Early Triassic of arctic Russia. It is known from a fairly complete skeleton discovered in a borehole on Kolguyev Island, though damage to the specimen and loss of certain bones has complicated study of the genus. Boreopricea shared many similarities with various other archosauromorphs, making its classification controversial. Various studies have considered it a close relative of Prolacerta, tanystropheids, both, or neither. Boreopricea is unique among early archosauromorphs due to possessing contact between the jugal and squamosal bones at the rear half of the skull.
Postparietals are cranial bones present in fish and many tetrapods. Although initially a pair of bones, many lineages possess postparietals which were fused into a single bone. The postparietals were dermal bones situated along the midline of the skull, behind the parietal bones. They formed part of the rear edge of the skull roof, and the lateral edge of each postparietal often contacts the tabular and supratemporal bones. In fish, the postparietals are elongated, typically the largest components of the skull roof. Tetrapods possessed shorter postparietals, which were reduced further and shifted towards the braincase in amniotes. At several points in synapsid evolution, the postparietals fused to each other and the tabulars during embryological development. This fusion produces the interparietal bone, which is inherited by mammals. Postparietals are common in extinct amphibians and early reptiles. However, most living amphibians and living reptiles lack postparietal bones, with a few exceptions.
Thanatotheristes is a genus of tyrannosaurid dinosaur from the Late Cretaceous of Laramidia, approximately 80.1-79.5 Ma. Thanatotheristes contains only one species, T. degrootorum. Fossils of this taxon are found in the Foremost Formation of Alberta, Canada, coexisting with medium-sized ceratopsids like Xenoceratops foremostensis and small pachycephalosaurids like Colepiocephale lambei.
Rugarhynchos is an extinct genus of doswelliid archosauriform from the Late Triassic of New Mexico. The only known species is Rugarhynchos sixmilensis. It was originally described as a species of Doswellia in 2012, before receiving its own genus in 2020. Rugarhynchos was a close relative of Doswellia and shared several features with it, such as the absence of an infratemporal fenestra and heavily textured skull bones. However, it could also be distinguished by many unique characteristics, such as a thick diagonal ridge on the side of the snout, blunt spikes on its osteoderms, and a complex suture between the quadratojugal, squamosal, and jugal. Non-metric multidimensional scaling and tooth morphology suggest that Rugarhynchos had a general skull anatomy convergent with some crocodyliforms, spinosaurids, and phytosaurs. However, its snout was somewhat less elongated than those other reptiles.
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