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The angular is a large bone in the lower jaw (mandible) of amphibians and reptiles (birds included), which is connected to all other lower jaw bones: the dentary (which is the entire lower jaw in mammals), the splenial, the suprangular, and the articular. It is homologous to the tympanic bone in mammals, due to the incorporation of several jaw bones into the mammalian middle ear early in mammal evolution.
In therapsids (mammal ancestors and their kin), the lower jaw is made up of the dentary (the mandible in mammals) and a group of smaller "postdentary" bones near the jaw joint. As the dentary increased in size over million of years, two of these postdentary bones, the articular and angular, became increasingly reduced and the dentary eventually made direct contact with the upper jaw. These postdentary bones, even before their articular function was lost, probably transmitted sound vibrations to the stapes and, in some therapsids, a bent plate that might have supported a membrane capable of detecting vibrations developed on the angular. Eventually, it developed into the ectotympanic ring which supports the tympanic membrane in the ears of modern mammals. [1]
The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea.
The ossicles are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sound vibrations sent from the ear drum to the fluid-filled labyrinth (cochlea). The absence of the auditory ossicles would constitute a moderate-to-severe hearing loss. The term "ossicle" literally means "tiny bone". Though the term may refer to any small bone throughout the body, it typically refers to the malleus, incus, and stapes of the middle ear.
Synapsida is one of the two major clades of vertebrate animals in the group Amniota, the other being the Sauropsida. The synapsids were the dominant land animals in the late Paleozoic and early Mesozoic, but the only group that survived into the Cenozoic are mammals. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals.
In anatomy, the temporomandibular joints (TMJ) are the two joints connecting the jawbone to the skull. It is a bilateral synovial articulation between the temporal bone of the skull above and the condylar process of mandible below; it is from these bones that its name is derived. The joints are unique in their bilateral function, being connected via the mandible.
The jaws are a pair of opposable articulated structures at the entrance of the mouth, typically used for grasping and manipulating food. The term jaws is also broadly applied to the whole of the structures constituting the vault of the mouth and serving to open and close it and is part of the body plan of humans and most animals.
Probainognathus meaning “progressive jaw” is an extinct genus of cynodonts that lived around 235 to 221.5 million years ago, during the Late Triassic in what is now Argentina. Together with the genus Bonacynodon from Brazil, Probainognathus forms the family Probainognathidae. Probainognathus was a relatively small, carnivorous or insectivorous cynodont. Like all cynodonts, it was a relative of mammals, and it possessed several mammal-like features. Like some other cynodonts, Probainognathus had a double jaw joint, which not only included the quadrate and articular bones like in more basal synapsids, but also the squamosal and surangular bones. A joint between the dentary and squamosal bones, as seen in modern mammals, was however absent in Probainognathus.
The articular bone is part of the lower jaw of most vertebrates, including most jawed fish, amphibians, birds and various kinds of reptiles, as well as ancestral mammals.
In humans, the cartilaginous bar of the mandibular arch is formed by what are known as Meckel's cartilages also known as Meckelian cartilages; above this the incus and malleus are developed. Meckel's cartilage arises from the first pharyngeal arch.
The tympanic part of the temporal bone is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the ear canal.
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone.
The evolution of mammalian auditory ossicles was an evolutionary process that resulted in the formation of the bones of the mammalian middle ear. These bones, or ossicles, are a defining characteristic of all mammals. The event is well-documented and important as a demonstration of transitional forms and exaptation, the re-purposing of existing structures during evolution.
Eodicynodon is an extinct genus of dicynodont therapsids, a highly diverse group of herbivorous synapsids that were widespread during the middle-late Permian and early Triassic. As its name suggests, Eodicynodon is the oldest and most primitive dicynodont yet identified, ranging from the middle to late Permian and possessing a mix of ancestral anomodont/therapsid features and derived dicynodont synapomorphies.
Morganucodonta is an extinct order of basal Mammaliaformes, a group including crown-group mammals (Mammalia) and their close relatives. Their remains have been found in Southern Africa, Western Europe, North America, India and China. The morganucodontans were probably insectivorous and nocturnal, though like eutriconodonts some species attained large sizes and were carnivorous. Nocturnality is believed to have evolved in the earliest mammals in the Triassic as a specialisation that allowed them to exploit a safer, night-time niche, while most larger predators were likely to have been active during the day.
In jawed vertebrates, the mandible, lower jaw, or jawbone is a bone that makes up the lower – and typically more mobile – component of the mouth.
In the auditory system, the columella contributes to hearing in amphibians, reptiles and birds. The columella form thin, bony structures in the interior of the skull and serve the purpose of transmitting sounds from the eardrum. It is an evolutionary homolog of the stapes, one of the auditory ossicles in mammals.
Karenites is an extinct genus of therocephalian therapsids from the Late Permian of Russia. The only species is Karenites ornamentatus, named in 1995. Several fossil specimens are known from the town of Kotelnich in Kirov Oblast.
The postdentary trough is a skeletal feature seen in Mesozoic mammals. It is found on the inside of the lower jaw (dentary), at the back behind the molar teeth. It is the hollow in which the postdentary bones and Meckel's cartilage sit. These bones form the middle ear in later mammal groups ; they include the incus (quadrate), malleus (articular), ectotympanic (angular) and prearticular.
The malleus, or hammer, is a hammer-shaped small bone or ossicle of the middle ear. It connects with the incus, and is attached to the inner surface of the eardrum. The word is Latin for 'hammer' or 'mallet'. It transmits the sound vibrations from the eardrum to the incus (anvil).
Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.
Bocaconodon is an early mammaliaform genus that lived during the Pliensbachian of Mexico. The type and only species, Bocaconodon tamaulipensis, was named and described in 2008. It is known from a single specimen, a partial right dentary bone preserving two nearly complete molar teeth and the rear portion of a third molar. The specimen was found at the Huizachal Canyon locality, "a Pliensbachian floodplain siltstone in the La Boca Formation".