Eocursor Temporal range: Early Jurassic, | |
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Restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Genus: | † Eocursor Butler, Smith & Norman, 2007 |
Species: | †E. parvus |
Binomial name | |
†Eocursor parvus Butler, Smith & Norman, 2007 | |
Eocursor (meaning "dawn runner") is genus of basal ornithischian dinosaur that lived in what is now South Africa during the Early Jurassic. Remains of this animal have been found in the Upper Elliot Formation and it is among the most completely known early ornithischians, shedding new light on the origin of the group. [1]
The exact age of this taxon has been the subject of uncertainty. It was originally interprereted as living during the Late Triassic (Norian age), around 210 million years ago; [1] however, Olsen, Kent & Whiteside (2010) stated that there is no independent geochronological support for its assumed age, and the available data makes it impossible to conclusively determine whether Eocursor is of Triassic or Early Jurassic (potentially as young as Sinemurian) age. [2] Eocursor was subsequently interpreted as a taxon of Early Jurassic age by McPhee et al. (2017), who identified the specimen as having originated from the Upper (not Lower) Elliot Formation. [3]
Fossils of Eocursor were originally collected in 1993, but were not formally described until fourteen years later. The type species, Eocursor parvus, was described in 2007 by Richard J. Butler, Roger M. H. Smith, and David B. Norman. Eocursor was one of the earliest known ornithischians, and sheds some light on early dinosaur relationships because early dinosaurs are known from mostly incomplete skeletons. Eocursor is known from partial skeletal elements, including skull fragments, spinal elements, pelvis, long leg bones, and unusually large grasping hands.
Eocursor was a lightly built bipedal dinosaur with an estimated length of about 1 m (3 ft). The general aspect of the animal resembles that of the early Jurassic ornithischians such as Lesothosaurus and Scutellosaurus . Its large hands resembled those of the Heterodontosauridae, a clade of primitive ornithischians. The morphology of the triangular teeth, not unlike those of an iguana, suggests partial herbivory. The tibia was significantly longer than the femur, indicating it was a swift runner. [1]
In 1993, the holotype specimen of Eocursor (SAM-PK-K8025) was collected from the Upper Elliot Formation in Free State, South Africa. The skeleton, although only partially complete, is one of the best-preserved early ornithischian skeletons ever found. The fossils include a partial skeleton: portions of the skull, lower jaw, vertebrae, and limbs. [1] [3]
In June 2007, the fossils were formally described by an international team of paleontologists: Richard Butler of the Natural History Museum, London, and the University of Cambridge, David Norman of the University of Cambridge, and Roger M. H. Smith of the Iziko South African Museum. The fossils were named Eocursor parvus, from the Greek word eos (meaning "dawn"), and the Latin words cursor (meaning "runner") and parvus ("little"), "in reference to the early occurrence of this ornithischian, its apparent locomotory abilities and its small size." [1]
Eocursor was an early ornithischian, one of the first so-called "bird-hipped" dinosaurs, a group which would eventually give rise to animals such as Stegosaurus , Triceratops , and Iguanodon . Butler et al. consider Eocursor more primitive than Lesothosaurus and Heterodontosauridae, but more derived than the Pisanosaurus , as a basal ornithischian forming a sister clade of Genasauria. [1]
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.
Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.
Abrictosaurus is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic in what is now in parts of southern Africa such as Lesotho and South Africa. It was a bipedal herbivore or omnivore and was one of the most basal heterodontosaurids. It was approximately 1.2 metres (3.9 ft) long and weighed between 0.68 and 3 kilograms.
Scelidosaurus is a genus of herbivorous armoured ornithischian dinosaur from the Jurassic of England.
Agilisaurus is a genus of ornithischian dinosaur from the Middle Jurassic Period of what is now eastern Asia. It was about 3.5–4 ft long, 2 ft in height and 40 kg in weight.
Echinodon is a genus of heterodontosaurid dinosaur that lived during the earliest Cretaceous of southern England and possibly western France in the Berriasian epoch. The first specimens were jaw bones named Echinodon becklesii by Sir Richard Owen in 1861, and since their original description only additional teeth have been discovered. The specific name honours collector Samuel Beckles who discovered the material of Echinodon and many other taxa from across England, while the genus name translates as "prickly tooth" in reference to the dental anatomy of the taxon.
Fabrosaurus is a dubious extinct genus of ornithischian dinosaur that lived during the Early Jurassic during the Hettangian to Sinemurian stages 199 - 189 mya. It was originally placed within the now obsolete family Fabrosauridae.
Heterodontosaurus is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic, 200–190 million years ago. Its only known member species, Heterodontosaurus tucki, was named in 1962 based on a skull discovered in South Africa. The genus name means "different toothed lizard", in reference to its unusual, heterodont dentition; the specific name honours G. C. Tuck, who supported the discoverers. Further specimens have since been found, including an almost complete skeleton in 1966.
Neornithischia is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs. Neornithischians include a variety of basal forms historically known as "hypsilophodonts", including the Parksosauridae; in addition, there are derived forms classified in the groups Marginocephalia and Ornithopoda. The former includes clades Pachycephalosauria and Ceratopsia, while the latter typically includes Hypsilophodon and the more derived Iguanodontia.
Pisanosaurus is an extinct genus of early dinosauriform, likely an ornithischian or silesaurid, from the Late Triassic of Argentina. It was a small, lightly built, ground-dwelling herbivore, that could grow up to an estimated 1 m (3.3 ft) long. Only one species, the type, Pisanosaurus mertii, is known, based on a single partial skeleton discovered in the Ischigualasto Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina. This part of the formation has been dated to the late Carnian, approximately 229 million years ago.
Lycorhinus is a genus of heterodontosaurid ornithischian dinosaur from the Early Jurassic strata of the Elliot Formation located in the Cape Province, South Africa.
Heterodontosauridae is a family of ornithischian dinosaurs that were likely among the most basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly found to be primitive, outside of the group Genasauria. Although their fossils are relatively rare and their group small in numbers, they have been found on all continents except Australia and Antarctica, with a range spanning the Early Jurassic to the Early Cretaceous.
Dracovenator is a genus of neotheropod dinosaur that lived approximately 201 to 199 million years ago during the early part of the Jurassic Period in what is now South Africa. Dracovenator was a medium-sized, moderately-built, ground-dwelling, bipedal carnivore, that could grow up to an estimated 5.5–6.5 metres (18–21 ft) in length and 250 kilograms (550 lb) in body mass. Its type specimen was based on only a partial skull that was recovered.
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town.
Fruitadens is a genus of heterodontosaurid dinosaur. The name means "Fruita teeth", in reference to Fruita, Colorado (USA), where its fossils were first found. It is known from partial skulls and skeletons from at least four individuals of differing biological ages, found in Tithonian rocks of the Morrison Formation in Colorado. Fruitadens is one of the smallest known ornithischian dinosaur, with young adults estimated at 65 to 75 cm in length and 0.5 to 0.75 kg in weight. It is interpreted as an omnivore and represents one of the latest-surviving heterodontosaurids.
Diodorus is a genus of silesaurid dinosauromorph that lived during the Late Triassic in what is now Morocco. Fossils were discovered in the Timezgadiouine Formation of the Argana Basin, and were used to name the new genus and species Diodorus scytobrachion. The genus name honors the mythological king Diodorus and the ancient historian Diodorus Siculus; the specific name is ancient Greek for 'leathery arm' and also honors the mythographer Dionysius Scytobrachion. The holotype specimen is a partial dentary bone (front of the lower jaw), and assigned specimens include isolated teeth, two humeri (upper arm bones), a metatarsal (foot bone), and femur (thigh bone).
Pegomastax is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic of South Africa. The only known specimen was discovered in a 1966–1967 expedition in Transkei District of Cape Province, but was not described until 2012 when Paul Sereno named it as the new taxon Pegomastax africana. The genus name is derived from the Greek for "strong jaw", and the species name describes the provenance of Africa; it was originally spelled africanus, was corrected to africana to align with the gender of the genus name.
Megapnosaurus is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 2.2 m (7.2 ft) long and weigh up to 13 kg (29 lb).
Laquintasaura is a genus of Venezuelan ornithischian dinosaur containing only the type species Laquintasaura venezuelae. The species was the first dinosaur to have been identified from Venezuela. It is known from extensive remains, all from a single bonebed locality which has been sampled for specimen blocks over the course of several expeditions, largely led by Marcelo R Sánchez-Villagra. A small animal, it is known for its distinct dental anatomy and for being one of the earliest and most primitive ornithischians in the fossil record. Taxonomic uncertainty has led to conflicting hypotheses that it is either at the base of Ornithischia or at the base of the subgroup Thyreophora. In either model, its dating to around 200 million years ago at the start of the Jurassic, existence in equatorial latitudes, and primitive nature make it a key insight into early ornithischian evolution. It is thought that Laquintasaura would have lived in groups and had a possible omnivorous diet, living on a seasonal alluvial plain and being preyed upon by the contemporary Tachiraptor.