Ajkaceratops Temporal range: Santonian ~ | |
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Holotype proposed rostral fused with premaxillae | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Ceratopsia |
Genus: | † Ajkaceratops Ősi et al., 2010 |
Species: | †A. kozmai |
Binomial name | |
†Ajkaceratops kozmai Ősi et al., 2010 | |
Ajkaceratops (pronounced "oi-ka-sera-tops") is a genus of ornithischian dinosaur described in 2010. It lived during the Late Cretaceous in the western Tethyan archipelago, in what is now Europe. The type species, A. kozmai, [1] was originally described as a ceratopsian most closely related to forms in east Asia, from where its ancestors may have migrated by island-hopping. Later research however has questioned this assignment, and treats Ajkaceratops as an ornithischian of unresolved affinity.
The holotype, cataloged as MTM V2009.192.1, consists only of a few skull fragments, including snout with proposed rostral bone, fused premaxillae, and maxillae fragments (beak and jaw fragments). These fossils are kept in the Hungarian Natural History Museum, in Budapest. The generic name, Ajkaceratops, honors Ajka, a town in Hungary where the fossils were first discovered, combined with the given greek nomination ceratops , meaning "horned face". The specific name, "kozmai", honors Károly Kozma. [1]
Although the fossils are fragmentary, the paper describing Ajkaceratops estimated a body length of 1 m (3.3 ft). [1] Other material includes four predentary bones, cataloged as MTM V2009.193.1, V2009.194.1, V2009.195.1, and V2009.196.1; these are also believed to have belonged to Ajkaceratops, although they are proportionately smaller, and probably came from other individuals of the genus. [1]
When first described, the fossils were stated to most closely resemble those of the Asian protoceratopsid Bagaceratops . Those similarities indicate Ajkaceratops is a ceratopsian related to the protoceratopsids, but more primitive than the Zuniceratops and the Ceratopsidae. [1]
However, more recent research has questioned this referral. The proposed rostral bone of Ajkaceratops differs markedly from the rostral morphology of other ceratopsians, possessing a sub-circular cross section, pitted texture instead of grooves, and a lack of lateral slicing edges. Furthermore, the proposed rostral bone appears to have fully fused with the premaxilla, a feature that is only seen in the ontogenetically oldest members of derived ceratopsids. This has cast doubt as to whether Ajkaceratops possesses a rostral bone in the first place, a defining feature of the ceratopsian group, and instead the hooked beak represents a highly derived, hypertrophied premaxilla. While there is the possibility Ajkaceratops is a highly unique ceratopsian, it is currently referred to as an enigmatic ornithischian of uncertain affinities. [2]
The fossils of Ajkaceratops were discovered in the Csehbánya Formation, which is interpreted as a floodplain and channel deposit formed by variegated clay, silt with interbedded grey and brown sand, and sandstone beds. This strata dates to the Santonian stage, around 86 to 84 million years ago. Ajkaceratops shared its environment with other dinosaurs such as Mochlodon , the nodosaurid ankylosaurs, other non-avian theropods and enantiornithine birds, as well as eusuchian crocodiles, azhdarchid pterosaurs, bothremydid turtles and teiid lizards. [3]
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.
Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.
Protoceratops is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus Protoceratops includes two species: P. andrewsi and the larger P. hellenikorhinus. The former was described in 1923 with fossils from the Mongolian Djadokhta Formation, and the latter in 2001 with fossils from the Chinese Bayan Mandahu Formation. Protoceratops was initially believed to be an ancestor of ankylosaurians and larger ceratopsians, such as Triceratops and relatives, until the discoveries of other protoceratopsids. Populations of P. andrewsi may have evolved into Bagaceratops through anagenesis.
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.
Claosaurus is a genus of hadrosauroid dinosaur that lived during the Late Cretaceous Period (Santonian-Campanian).
Echinodon is a genus of heterodontosaurid dinosaur that lived during the earliest Cretaceous of southern England and possibly western France in the Berriasian epoch. The first specimens were jaw bones named Echinodon becklesii by Sir Richard Owen in 1861, and since their original description only additional teeth have been discovered. The specific name honours collector Samuel Beckles who discovered the material of Echinodon and many other taxa from across England, while the genus name translates as "prickly tooth" in reference to the dental anatomy of the taxon.
Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most specimens found in China and Mongolia. As ceratopsians, protoceratopsids were herbivorous, with constantly replacing tooth batteries made for slicing through plants and a hooked beak for grabbing them. Protoceratopsids were small ceratopsians around 1-2.5 m in length. Their bony frill and horns were much smaller than more derived members of Ceratopsia, such as ceratopsids.
Arrhinoceratops is a genus of herbivorous ceratopsian dinosaur. The name was coined as its original describer concluded it was special because the nose-horn was not a separate bone, however further analysis revealed this was based on a misunderstanding. It lived during the latest Campanian/earliest Maastrichtian stage of the Late Cretaceous, predating its famous relative Triceratops by a few million years, although it was contemporary with Anchiceratops. Its remains have been found in Canada.
Heterodontosaurus is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic, 200–190 million years ago. Its only known member species, Heterodontosaurus tucki, was named in 1962 based on a skull discovered in South Africa. The genus name means "different toothed lizard", in reference to its unusual, heterodont dentition; the specific name honours G. C. Tuck, who supported the discoverers. Further specimens have since been found, including an almost complete skeleton in 1966.
Hungarosaurus tormai, is a herbivorous nodosaurid ankylosaur from the Upper Cretaceous (Santonian) Csehbánya Formation of the Bakony Mountains of western Hungary. It is the most completely known ankylosaur from the Cretaceous of Europe.
Breviceratops is a genus of protoceratopsid dinosaur that lived during the Late Cretaceous in what is now the Barun Goyot Formation, Mongolia.
Zalmoxes is a genus of rhabdodontid ornithopod dinosaur from the Maastrichtian age of the Late Cretaceous in what is now Romania. The genus is known from specimens first named as the species Mochlodon robustum in 1899 by Franz Nopcsa before being reclassified as Rhabdodon robustum by him in 1915. In 1990, this name was corrected to Rhabdodon robustus by George Olshevsky and, in 2003, the species was once more reclassified as the type species Zalmoxes robustus. Zalmoxes refers to the Dacian deity Zalmoxis and robustus refers to the robustness of the remains. Also in 2003, another species was named, Zalmoxes shqiperorum, named for the Albanian name for Albanians.
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Bakonydraco is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western Hungary.
Rhabdodontidae is a family of herbivorous iguanodontian ornithopod dinosaurs whose earliest stem members appeared in the middle of the Lower Cretaceous. The oldest dated fossils of these stem members were found in the Barremian Castrillo de la Reina Formation of Spain, dating to approximately 129.4 to 125.0 million years ago. With their deep skulls and jaws, Rhabdodontids were similar to large, robust iguanodonts. The family was first proposed by David B. Weishampel and colleagues in 2003. Rhabdodontid fossils have been mainly found in Europe in formations dating to the Late Cretaceous.
The Csehbánya Formation is a geological formation in the Transdanubian Mountains of Veszprém County, Hungary. The formation dates to the Late Santonian of the Late Cretaceous. It represents a floodplain environment as opposed to the swampy lacustrine environment of the simultaneous Ajka Coal Formation, though there is complete overlap in terms of fauna. It underlies the Jákó Marl Formation, and laterally transitions to the Ajka Coal Formation.
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This timeline of ceratopsian research is a chronological listing of events in the history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bony frills, and, later, spectacular horns. The first scientifically documented ceratopsian fossils were described by Edward Drinker Cope starting in the 1870s; however, the remains were poorly preserved and their true nature was not recognized. Over the next several decades, Cope named several such genera and species. Cope's hated rival, Othniel Charles Marsh, also described ceratopsian remains. In 1887, Marsh mistook a Triceratops horn for one belonging to a new species of prehistoric Bison. Marsh also named the eponymous genus Ceratops in 1888. The next year, he named the most famous ceratopsian, Triceratops horridus. It was the discovery of Triceratops that illuminated the ceratopsian body plan, and he formally named the Ceratopsia in 1890.
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