Chaoyangsauridae Temporal range: Late Jurassic, | |
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Compared close-up of the four so far known chaoyangsaurid genera, with pink representing known parts of skulls | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ceratopsia |
Family: | † Chaoyangsauridae Zhao, Cheng, & Xu, 1999 |
Type species | |
† Chaoyangsaurus youngi Zhao, Cheng, & Xu, 1999 | |
Genera | |
Chaoyangsauridae is a family of ceratopsian dinosaurs. They are among the earliest known marginocephalian dinosaurs, with remains dating to about 160 million years ago, during the Late Jurassic period. Members of this group had sharp beaks for snipping off leaves to eat, and a very small frill.
This family is officially defined in the PhyloCode as "the largest clade containing Chaoyangosaurus youngi , but not Psittacosaurus mongoliensis and Triceratops horridus ". [1]
Four dinosaur genera, Chaoyangsaurus , Xuanhuaceratops , Yinlong and Hualianceratops , are usually considered to belong to the Chaoyangsauridae. [2] [3] [4] In 2020, Yu et al. classified Stenopelix as a chaoyangsaurid. [5] All five animals are more primitive (or basal) than both Psittacosaurus and neoceratopsians.
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period, although ancestral forms lived earlier, in the Late Jurassic of Asia. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.
Marginocephalia is a clade of ornithischian dinosaurs that is characterized by a bony shelf or margin at the back of the skull. These fringes were likely used for display. This clade was officially defined in the PhyloCode by Daniel Madzia and colleagues in 2021 as "the smallest clade containing Ceratops montanus, Pachycephalosaurus wyomingensis, and Triceratops horridus". There are two clades included in Marginocephalia: the thick-skulled Pachycephalosauria and the horned Ceratopsia. All members of Marginocephalia were primarily herbivores. They basally used gastroliths to aid in digestion of tough plant matter until they convergently evolved tooth batteries in Neoceratopsia and Pachycephalosauria. Marginocephalia first evolved in the Jurassic Period and became more common in the Cretaceous. They are basally small facultative quadrupeds while derived members of the group are large obligate quadrupeds. Primitive marginocephalians are found in Asia, but the group migrated upwards into North America.
Psittacosaurus is a genus of extinct ceratopsian dinosaur from the Early Cretaceous of what is now Asia, existing between 125 and 105 million years ago. It is notable for being the most species-rich non-avian dinosaur genus. Up to 12 species are known, from across China, Mongolia, Russia, and Thailand. The species of Psittacosaurus were obligate bipeds at adulthood, with a high skull and a robust beak. One individual was found preserved with long filaments on the tail, similar to those of Tianyulong. Psittacosaurus probably had complex behaviours, based on the proportions and relative size of the brain. It may have been active for short periods of time during the day and night, and had well-developed senses of smell and vision.
Chaoyangsaurus is a marginocephalian dinosaur from the Late Jurassic of China. It has been dated to between 150.8 and 145.5 million years ago. Chaoyangsaurus belonged to the Ceratopsia. Chaoyangsaurus, like all ceratopsians, was primarily a herbivore.
Stenopelix is a genus of probable basal ceratopsian from the Early Cretaceous of Germany. It lived in the late Berriasian Stage of the Cretaceous period, approximately 140 myr ago. The genus is based on a partial skeleton lacking the skull, and its classification is based on characteristics of the hips.
Xuanhuaceratops is a genus of dinosaur from the Late Jurassic Period. The genus was in 2006 dated to the Tithonian, 150.8-145.5 million years ago. A member of the family Chaoyangsauridae, it was one of the earliest ceratopsians. The fossils were found in the Houcheng Formation of Hebei Province, northeastern China.
Yinlong is a genus of basal ceratopsian dinosaur from the Late Jurassic Period of China. By far the earliest known ceratopsian, it was a small, primarily bipedal herbivore.
Xu Xing is a Chinese paleontologist who has named more dinosaurs than any other living paleontologist. Such dinosaurs include the Jurassic ceratopsian Yinlong, the Jurassic tyrannosauroid Guanlong, the large oviraptorosaur Gigantoraptor, and the troodontid Mei.
The Shishugou Formation is a geological formation in Xinjiang, China.
Albalophosaurus is a genus of marginocephalian ornithischian dinosaur that lived in Japan during the Early Cretaceous. The type species is Albalophosaurus yamaguchiorum.
Haplocheirus is an extinct genus of theropod dinosaur from the Middle Jurassic Shishugou Formation of Xinjiang in China. It is generally considered to be an alvarezsauroid, although some researchers have questioned this assignment. The genus contains a single species, H. sollers, which is known from a mostly complete skeleton including the skull.
Koreaceratops is a genus of basal ceratopsian dinosaur discovered in Albian-age Lower Cretaceous rocks of South Korea.
Aorun is a genus of carnivorous theropod dinosaur first discovered in 2006, with its scientific description published in 2013. It is possibly one of the oldest known coelurosaurian dinosaurs and is estimated to have lived ~161.6 million years ago during the Late Jurassic Period, though some researchers consider it to be a carnosaurian instead.
This timeline of ceratopsian research is a chronological listing of events in the history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bony frills, and, later, spectacular horns. The first scientifically documented ceratopsian fossils were described by Edward Drinker Cope starting in the 1870s; however, the remains were poorly preserved and their true nature was not recognized. Over the next several decades, Cope named several such genera and species. Cope's hated rival, Othniel Charles Marsh, also described ceratopsian remains. In 1887, Marsh mistook a Triceratops horn for one belonging to a new species of prehistoric Bison. Marsh also named the eponymous genus Ceratops in 1888. The next year, he named the most famous ceratopsian, Triceratops horridus. It was the discovery of Triceratops that illuminated the ceratopsian body plan, and he formally named the Ceratopsia in 1890.
Mosaiceratops is a genus of ceratopsian, described by Zheng, Jin & Xu in 2015 and found in the Xiaguan Formation of Neixiang County. Mosaiceratops lived in the upper Cretaceous in what is now the Henan Province of China.
Hualianceratops is a genus of herbivorous ceratopsian dinosaur that lived about 160 million years ago in the Late Jurassic epoch in what is now western China. The single species, H. wucaiwanensis was described in 2015. Its size has been compared to that of a spaniel.
Isaberrysaura is a genus of stegosaurian ornithischian dinosaur from the Middle Jurassic Los Molles Formation of Patagonia, Argentina. The genus contains a single species, I. mollensis, described by Salgado et al. in 2017 from a single specimen. Although initially classified as a basal neornithischian, subsequent analysis has allied it with the Stegosauria; the morphology of its skull resembles those of other members of the group.
Beg is a genus of neoceratopsian dinosaur from the early Cretaceous period of Mongolia. The genus contains a single species, Beg tse, known from a partial skull and very fragmentary postcrania. Beg represents the most basal neoceratopsian currently known.