Unescoceratops Temporal range: Late Cretaceous, | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Suborder: | † Ceratopsia |
Family: | † Leptoceratopsidae |
Genus: | † Unescoceratops Ryan et al., 2012 [1] |
Species: | †U. koppelhusae |
Binomial name | |
†Unescoceratops koppelhusae Ryan et al., 2012 [1] | |
Unescoceratops is a genus of leptoceratopsid ceratopsian dinosaurs known from the Late Cretaceous of Alberta, southern Canada. It contains a single species, Unescoceratops koppelhusae. [1]
Unescoceratops is known only from the holotype specimen TMP 95.12.6, a partial left dentary. The fossil was collected in 1995 in the Black Coulee locality (formerly Deadhorse Coulee), near the Writing-on-Stone Provincial Park, from the Dinosaur Park Formation, dating to the late Campanian stage of the Late Cretaceous period, about 76.5-75 million years ago. The specimen was regarded as too incomplete to identify, and was shelved for several years. It was believed to be a Leptoceratops specimen at the time. A cladistic analysis done by Michael Ryan (of The Cleveland Museum of Natural History) and David Evans (of the Royal Ontario Museum) found it to be among the most advanced leptoceratopsid genera. [1] [2] [3]
Unescoceratops is thought to have been between one and two meters long and less than 91 kilograms. Its teeth were the roundest of all leptoceratopsids. [3]
Mallon et al. (2013) examined herbivore coexistence on the island continent of Laramidia, during the Late Cretaceous. It was concluded that small ornithischians like Unescoceratops were generally restricted to feeding on vegetation at, or below the height of 1 meter. [4]
The genus Unescoceratops was first described by Michael J. Ryan, David C. Evans, Philip J. Currie, Caleb M. Brown and Don Brinkman in 2012 (though it had been published earlier by Ryan and Currie in 1995 with no name) and the type species is Unescoceratops koppelhusae. [1] The generic name means "UNESCO horned face, in reference to the fossil being found in an area designated as a World Heritage Site by the organization. "'Dinosaurprovincialparkaceratops' was too long," explained Ryan. [2] The specific epithet is named after Currie's wife, palynologist Eva Koppelhus. [4]
Unescoceratops has been classified as a leptoceratopsid and the sister taxa to Gryphoceratops , [1] which was described in the same paper. [5] These two are the most derived leptoceratopsids known, despite Gryphoceratops also being the oldest leptoceratopsid. [1]
The cladogram below represents the cladistic relationship of Unescoceratops compared to other leptoceratopsids as reconstructed from the phylogenetic analysis. [1]
Leptoceratopsidae |
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Chasmosaurus is a genus of ceratopsid dinosaur from the Late Cretaceous Period in North America. Its given name means 'opening lizard', referring to the large openings (fenestrae) in its frill. With a length of 4.3–4.8 metres (14.1–15.7 ft) and a weight of 1.5–2 tonnes —or anywhere from 2,200 to nearly 5,000 lbs., give or take—Chasmosaurus was of a slightly smaller to ‘average’ size, especially when compared to larger ceratopsians. The Chasmosaurs were similar, in overall build and weight, to a white rhinoceros or an Indian rhinoceros; just like rhinos, and all other ceratopsians, they were purely herbivorous, needing to consume around 54 kilograms, or 120 lbs., of plant matter each day.
Styracosaurus is an extinct genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of North America. It had four to six long parietal spikes extending from its neck frill, a smaller jugal horn on each of its cheeks, and a single horn protruding from its nose, which may have been up to 60 centimeters long and 15 centimeters wide. The function or functions of the horns and frills have been debated for many years.
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Montanoceratops is an extinct genus of small ceratopsian dinosaur that lived approximately 70 million years ago during the latter part of the Cretaceous Period in what is now Montana and Alberta. Montanoceratops was a small sized, moderately-built, ground-dwelling, quadrupedal herbivore, that could grow up to an estimated 2.5 m (8.2 ft) in length and 170 kg (370 lb) in body mass.
Asiaceratops is a genus of herbivorous ceratopsian dinosaur. It lived during the Early-Late Cretaceous. The type species, A. salsopaludalis is known from Uzbekistan, while A. sulcidens is known from China and Mongolia.
Zapsalis is a genus of dromaeosaurine theropod dinosaurs. It is a tooth taxon, often considered dubious because of the fragmentary nature of the fossils, which include teeth but no other remains.
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Prenoceratops, is a genus of herbivorous ceratopsian dinosaur from the Late Cretaceous Period. It was a relatively small dinosaur, reaching 1.3 m (4.3 ft) in length and 20 kg (44 lb) in body mass. Its fossils have been found in the upper Two Medicine Formation in the present-day U.S. state of Montana, in Campanian age rock layers that have been dated to 74.3 million years ago. Fossils were also found in the Oldman Formation in the modern day Canadian province of Alberta, dating to around 77 million years ago.
Medusaceratops is an extinct genus of centrosaurine ceratopsian dinosaur known from the Late Cretaceous Judith River Formation of Montana, northern United States. It contains a single species, Medusaceratops lokii.
Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller.
The Milk River Formation is a sandstone-dominated stratigraphic unit of the Western Canada Sedimentary Basin in southern Alberta, Canada. It was deposited in near-shore to coastal environments during Late Cretaceous time. Based on uranium-lead dating, palynology and stratigraphic relationships, deposition occurred between ~84.1 and 83.6 Ma.
Gryphoceratops is an extinct genus of leptoceratopsid ceratopsian dinosaur known from the Late Cretaceous of Alberta, southern Canada.
Spinops is an extinct genus of centrosaurine ceratopsian dinosaur from the Late Cretaceous of Alberta, southern Canada. It was a medium-sized ceratopsian, reaching 4.5 metres (15 ft) in length and 1.3 metric tons in body mass.
Coronosaurus is a genus of centrosaurine ceratopsian dinosaurs which lived in the Late Cretaceous, in the middle Campanian stage. Its remains, two bone beds, were discovered by Phillip J. Currie in the Oldman Formation of Alberta, Canada, and its type and only species, Coronosaurus brinkmani, was first described in 2005, as a new species within the genus Centrosaurus. Later studies questioned the presence of a direct relationship, and in 2012 it was named as a separate genus. Coronosaurus means "crowned lizard", coming from "corona", Latin for crown, and "sauros", Greek for lizard; this name refers to the unique, crown-like shape of the horns on the top of its frill.
This timeline of ceratopsian research is a chronological listing of events in the history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bony frills, and, later, spectacular horns. The first scientifically documented ceratopsian fossils were described by Edward Drinker Cope starting in the 1870s; however, the remains were poorly preserved and their true nature was not recognized. Over the next several decades, Cope named several such genera and species. Cope's hated rival, Othniel Charles Marsh, also described ceratopsian remains. In 1887, Marsh mistook a Triceratops horn for one belonging to a new species of prehistoric Bison. Marsh also named the eponymous genus Ceratops in 1888. The next year, he named the most famous ceratopsian, Triceratops horridus. It was the discovery of Triceratops that illuminated the ceratopsian body plan, and he formally named the Ceratopsia in 1890.
Ischioceratops is an extinct genus of small herbivorous ceratopsian dinosaur that lived approximately 69 million years ago during the latter part of the Cretaceous Period in what is now China.
David Christopher Evans is a Canadian palaeontologist and evolutionary biologist who specializes in the evolution and paleobiology of Cretaceous dinosaurs in western North America. He received his B.Sc. from the University of British Columbia and his Ph.D. from the University of Toronto. He is a fellow of the Royal Canadian Geographical Society (RCGS) and a member of the Royal Society of Canada and currently serves as the Senior Curator and Temerty Chair of Vertebrate Paleontology at the Royal Ontario Museum in Toronto, Canada. He is also a faculty member in the Department of Ecology & Evolutionary Biology at the University of Toronto. Evans is particularly renowned for his work on the paleobiology of hadrosaur ("duck-billed") dinosaurs and has conducted international research on a wide variety of paleontological topics.
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