Stenopelix Temporal range: Late Berriasian, ~ | |
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Latex casts of the holotype slabs and interpretive drawings | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ceratopsia |
Family: | † Chaoyangsauridae |
Genus: | † Stenopelix Meyer, 1857 |
Species: | †S. valdensis |
Binomial name | |
†Stenopelix valdensis Meyer, 1857 | |
Stenopelix (meaning "narrow pelvis") is a genus of small marginocephalian dinosaur, possibly a basal ceratopsian, from the Early Cretaceous of Germany. It lived in the late Berriasian Stage of the Cretaceous period, approximately 140 myr ago. [1] The genus is based on a partial skeleton lacking the skull, and its classification is based on characteristics of the hips.
In 1855, in a sandstone quarry near Bückeburg on the Harrl, a fossil was found of a small dinosaur. Most of its bones were in a poor condition and removed on preparation, leaving two sets of hollow impressions on the plate and counterplate. The two plates do not overlap completely. The hollows, serving as a natural mold, have since been used to produce several casts in gypsum and latex to facilitate the study of the specimen. It was originally part of the collection of Max Ballerstedt preserved in the Bückeburg Gymnasium Adolfinum [2] but was in 1976 moved to the Georg-August-Universität Göttingen where it now resides in the collection of the Geowissenschaftliches Zentrum der Universität Göttingen.
In 1857, based on this fossil, Christian Erich Hermann von Meyer named the type species Stenopelix valdensis. [3] The generic name is derived from Greek stenos, "narrow", and pelyx, "pelvis". The specific name refers to the Wealden Formation. The holotype, GZG 741/2 (earlier GPI 741–1, 2), found in the Obernkirchen Sandstein Formation, consists of the impressions of an almost complete skeleton, lacking the skull and the neck.
Stenopelix was a small herbivorous animal, reaching 1.4 m (4.6 ft) in length and 10 kg (22 lb) in body mass. [4] The preserved rump and tail have a combined length of just 97 centimetres; the femur is fourteen centimetres long. The species can be distinguished by several details of the pelvis. The shaft part of the ilium uniformly tapers ending in a rounded point. The shaft of the ischium is thickest in the middle and there shows a distinctive kink.
The classification of Stenopelix is controversial and has ever been problematic because of the lacking skull. Prior to the 1960s, it was often assigned to some ornithopod group. In 1974 Teresa Maryańska suggested it to be a pachycephalosaur, one of the oldest known, due to the apparent exclusion of the pubis from the acetabulum, and the presence of strong caudal ribs. Peter Galton in 1982 showed that the "pubis" was actually part of the acetabulum, and the so-called "caudal ribs" were sacral ribs. The curvature of the ischium and absence of an obturator foramen were not characteristics seen in other pachycephalosaurs. Galton concluded Stenopelix to be ceratopian. [5]
However, exact cladistic analyses by Paul Sereno have resulted in a position in the Pachycephalosauria. But paleontologists Richard J. Butler and Robert M. Sullivan nonetheless view the species as being Marginocephalia incertae sedis , rejecting the presumed synapomorphies with the Pachycephalosauria as incorrect identifications or lacking cogency because of a possible presence in ceratopsian groups. [6] A cladistic analysis performed by Butler et al. (2011) showed that Stenopelix is a basal member of the Ceratopsia, and its sister taxon is Yinlong . [7] Yu et al. (2020) classified Stenopelix as a chaoyangsaurid with Yinlong, Chaoyangsaurus, Xuanhuaceratops, Hualianceratops. [8] Fonseca et al. (2024) also recovered this genus as a chaoyangsaurid close to Yinlong. [9]
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.
Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.
Ornithopoda is a clade of ornithischian dinosaurs, called ornithopods. They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively small-sized, while advanced members of the subgroup Iguanodontia became quadrupedal and developed large body size. Their major evolutionary advantage was the progressive development of a chewing apparatus that became the most sophisticated ever developed by a non-avian dinosaur, rivaling that of modern mammals such as the domestic cow. They reached their apex of diversity and ecological dominance in the hadrosaurids, before they were wiped out by the Cretaceous–Paleogene extinction event along with all other non-avian dinosaurs. Members are known worldwide.
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.
Marginocephalia is a clade of ornithischian dinosaurs that is characterized by a bony shelf or margin at the back of the skull. These fringes were likely used for display. There are two clades included in Marginocephalia: the thick-skulled Pachycephalosauria and the horned Ceratopsia. All members of Marginocephalia were primarily herbivores. They basally used gastroliths to aid in digestion of tough plant matter until they convergently evolved tooth batteries in Neoceratopsia and Pachycephalosauria. Marginocephalia first evolved in the Jurassic Period and became more common in the Cretaceous. They are basally small facultative quadrupeds while derived members of the group are large obligate quadrupeds. Primitive marginocephalians are found in Asia, but the group migrated upwards into North America.
Pachycephalosauria is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephalosaurs lived during the Late Cretaceous Period, dating between about 85.8 and 66 million years ago. They are exclusive to the Northern Hemisphere, all of them being found in North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls. Skulls can be domed, flat, or wedge-shaped depending on the species, and are all heavily ossified. The domes were often surrounded by nodes and/or spikes. Partial skeletons have been found of several pachycephalosaur species, but to date no complete skeletons have been discovered. Often isolated skull fragments are the only bones that are found.
Echinodon is a genus of heterodontosaurid dinosaur that lived during the earliest Cretaceous of southern England and possibly western France in the Berriasian epoch. The first specimens were jaw bones named Echinodon becklesii by Sir Richard Owen in 1861, and since their original description only additional teeth have been discovered. The specific name honours collector Samuel Beckles who discovered the material of Echinodon and many other taxa from across England, while the genus name translates as "prickly tooth" in reference to the dental anatomy of the taxon.
Micropachycephalosaurus is an extinct genus of basal marginocephalian dinosaur containing only the type species, Micropachycephalosaurus hongtuyanensis. It lived in China during the Late Cretaceous (Campanian) and was found in the Jiangjunding Formation.
Yaverlandia is a genus of maniraptoran dinosaur. Known from a partial fossil skull found in Lower Cretaceous strata of the Wessex Formation on the Isle of Wight. it was described as the earliest known member of the pachycephalosaurid family, but research by Darren Naish shows it to have actually been a theropod, seemingly a maniraptoran. The type species is Y. bitholus.
Tylocephale is a genus of pachycephalosaurid dinosaur, a group of dome-headed, herbivorous ornithischians, that lived during the Late Campanian stage of the Late Cretaceous in what is now Mongolia. It is known from a partial skull and associated mandible that were unearthed in 1971 by a Polish-Mongolian Expedition to the Barun Goyot Formation of the Gobi Desert. The specimen was described in 1974 by Polish paleontologists Teresa Maryańska and Halszka Osmólska as a new genus and species.
Neornithischia is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs. Neornithischians include a variety of basal forms historically known as "hypsilophodonts", including the Parksosauridae; in addition, there are derived forms classified in the groups Marginocephalia and Ornithopoda. The former includes clades Pachycephalosauria and Ceratopsia, while the latter typically includes Hypsilophodon and the more derived Iguanodontia.
Graciliceratops is a genus of neoceratopsian dinosaurs that lived in Asia during the Late Cretaceous period.
Yinlong is a genus of basal ceratopsian dinosaur from the Late Jurassic Period of central Asia. By far the earliest known ceratopsian, it was a small, primarily bipedal herbivore.
Alaskacephale is an extinct genus of pachycephalosaurid, a group of dome-headed, herbivorous ornithischian dinosaurs, that lived during the Campanian stage of the Late Cretaceous period in what is now northern Alaska. The genus is one of the few known Arctic dinosaurs and was found in the Prince Creek Formation, which preserves a menagerie of fossils. The only known specimen, a squamosal bone, was found in 1999 and later described in 2005. However, Alaskacephale was not formally named until the next year.
Chaoyangsauridae is a family of ceratopsian dinosaurs. They are among the earliest known marginocephalian dinosaurs, with remains dating to about 160 million years ago, during the Late Jurassic period. Members of this group had sharp beaks for snipping off leaves to eat, and a very small frill.
Albalophosaurus is a genus of marginocephalian ornithischian dinosaur that lived in Japan during the Early Cretaceous. The type species is Albalophosaurus yamaguchiorum.
Koreaceratops is a genus of basal ceratopsian dinosaur discovered in Albian-age Lower Cretaceous rocks of South Korea.
This timeline of pachycephalosaur research is a chronological listing of events in the history of paleontology focused on the pachycephalosaurs, a group of dome-skulled herbivorous marginocephalian dinosaurs. One of the first major events related to the history of pachycephalosaur research actually regards the discovery of an unrelated dinosaur called Troodon, reported from the western United States by Joseph Leidy in 1856. The type specimen of Troodon was simply an unusual tooth, but the close resemblance between Troodon teeth and pachycephalosaur teeth would cause taxonomic confusion for over a century. This was resolved by Phil Currie in 1987, who realized that Troodon belonged to a group of bird-like carnivores then known as saurornithoidids, but since renamed Troodontidae after Troodon itself. The first scientifically documented true pachycephalosaur remains were discovered in Early Cretaceous rocks from England and named Stenopelix not long after Troodon was named in America. Other notable early finds include the well-known pachycephalosaur Stegoceras validum.
Mosaiceratops is a genus of ceratopsian, described by Zheng, Jin & Xu in 2015 and found in the Xiaguan Formation of Neixiang County. Mosaiceratops lived in the upper Cretaceous in what is now the Henan Province of China.
Beg is a genus of neoceratopsian dinosaur from the early Cretaceous period of Mongolia. The genus contains a single species, Beg tse, known from a partial skull and very fragmentary postcrania. Beg represents the most basal neoceratopsian currently known.