Brachyceratops Temporal range: Late Cretaceous, | |
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Holotype specimen USNM 7951 mounted in the Smithsonian | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ceratopsia |
Family: | † Ceratopsidae |
Subfamily: | † Centrosaurinae |
Genus: | † Brachyceratops Gilmore, 1914 |
Species: | †B. montanensis |
Binomial name | |
†Brachyceratops montanensis Gilmore, 1914 | |
Brachyceratops ('short horned face') is a dubious genus of ceratopsian dinosaur known only from partial juvenile specimens dating to the late Cretaceous Period of Montana, United States.
Brachyceratops has historically been known from juvenile remains, with one specimen having since been re-classified as Rubeosaurus ovatus . [1] [2]
Brachyceratops montanensis, the type species, was first discovered in the Two Medicine Formation (Campanian, about 74 million years old) on a Blackfoot Indian Reservation in Teton County in north-central Montana. [3] The original find was made in August 1913 by C. W. Gilmore [3] and his assistant John Floyd Strayer and was named and shortly described by Gilmore one year later. The generic name is derived from Greek : βραχύς, brachys, "short", Greek : κέρας, keras, "horn" and Greek : ὤψ, ops, "face", in reference to the short snout. The specific name refers to the provenance from Montana. [4]
All that was found were incomplete and jumbled remains of five juvenile individuals of about 1.5 m (5 feet) in length. [3] It has been speculated that these juveniles may have been nest mates that stayed together after hatching. [3] The holotype specimen is USNM 7951, a partial skull. The paratypes are USNM 7952, a snout, USNM 7953, a partial skeleton with skull and USNM 7957, a foot. The material is disarticulated but the preservation is excellent. [5] In 1917 Gilmore published a monograph on Brachyceratops in which a reconstruction of the skeleton as a whole was given. [6]
In 1939 Gilmore referred a larger subadult specimen, USNM 14765, to Brachyceratops. [7] All specimens are currently part of the collection of the Smithsonian Institution in Washington D.C., [3] where a skeletal restoration is mounted.
As Brachyceratops is known only from the remains of five juveniles — plus the subadult that Gilmore found about a mile from the original specimens —, it was long considered likely that these represented the immature forms of known centrosaurine ceratopsians, [3] [2] with Monoclonius often suggested as the likeliest candidate. By 2011 however, several studies had shown that the single specimen which was adult enough to be compared with related species, USNM 14765, represented the juvenile form of the later-named Rubeosaurus .
Among the five original specimens parts of three skulls were discovered, detached from their owner's body and fragmented. [3] Despite this, the skull showed that the animal had only small bumps over the eyes rather than full-grown horns like in the more famous ceratopsians such as Triceratops . [3] The nasal horn was thick and low, while its neck frill was moderately large. [3] Unfortunately the specimens were incomplete so it cannot be determined if there were parietal openings in the frill like some other ceratopsians possessed. [3]
Brachyceratops belonged to the Centrosaurinae, but its exact placement has been difficult to discover because it is known only from juvenile specimens. In 1997, Scott Sampson and colleagues re-examined Brachyceratops and noted that it is almost certainly the juvenile form of an already known centrosaurine dinosaur. However, because many features that distinguish ceratopsians from each other do not appear until adulthood, exactly which centrosaurine remained unknown, and Sampson et al. classified Brachyceratops as a nomen dubium, or dubious name. [8] In 2007, Michael J. Ryan and colleagues suggested that Brachyceratops was possibly the juvenile form of Styracosaurus ovatus, which has since been reclassified as Rubeosaurus . [1] A 2011 study supported this idea for the most mature specimen of Brachyceratops, USNM 14765, which shows one unique newly evolved feature (apomorphy) in common with Rubeosaurus to the exclusion of other centrosaurines. However, the same study suggested that because the holotype specimen of Brachyceratops is too incomplete and juvenile to preserve any determinable apomorphies, Brachyceratops must be considered a nomen dubium, and cannot be a senior synonym of Rubeosaurus. [9]
Dinosaurs that lived alongside Brachyceratops include the basal ornithopod Orodromeus , hadrosaurids (such as Hypacrosaurus , Maiasaura , and Prosaurolophus ), the centrosaurines Stellasaurus and Einiosaurus , the ankylosaurs Edmontonia and Euoplocephalus , the tyrannosaurid Daspletosaurus (which appears to have been a specialist of preying on ceratopsians), as well as the smaller theropods Bambiraptor , Chirostenotes , Troodon , and Avisaurus .
Torosaurus is a genus of herbivorous chasmosaurine ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous period, between 68 and 66 million years ago, though it is possible that the species range might extend to as far back as 69 million years ago. Fossils have been discovered across the Western Interior of North America, from as far north as Saskatchewan to as far south as Texas.
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.
Ceratopsidae is a family of ceratopsian dinosaurs including Triceratops, Centrosaurus, and Styracosaurus. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are known from western North America, which formed the island continent of Laramidia during most of the Late Cretaceous. Ceratopsids are characterized by beaks, rows of shearing teeth in the back of the jaw, elaborate nasal horns, and a thin parietal-squamosal shelf that extends back and up into a frill. The group is divided into two subfamilies—Chasmosaurinae and Centrosaurinae. The chasmosaurines are generally characterized by long, triangular frills and well-developed brow horns. The centrosaurines had well-developed nasal horns or nasal bosses, shorter and more rectangular frills, and elaborate spines on the back of the frill.
Nedoceratops is a controversial genus of ceratopsid dinosaur from the Late Cretaceous period Lance Formation of North America. It is known only from a single skull discovered in Wyoming. Its status is the subject of ongoing debate among paleontologists: some authors consider Nedoceratops a valid, distinct taxon, while others consider it to be an unusual specimen of Triceratops.
Styracosaurus is an extinct genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of North America. It had four to six long parietal spikes extending from its neck frill, a smaller jugal horn on each of its cheeks, and a single horn protruding from its nose, which may have been up to 60 centimeters long and 15 centimeters wide. The function or functions of the horns and frills have been debated for many years.
Einiosaurus is a genus of herbivorous centrosaurine ceratopsian dinosaur from the Upper Cretaceous of northwestern Montana. The name means 'bison lizard', in a combination of Blackfeet Indian eini and Latinized Ancient Greek sauros; the specific name (procurvicornis) means 'with a forward-curving horn' in Latin. Einiosaurus is medium-sized with an estimated body length at 4.5 metres (15 ft).
Achelousaurus is a genus of centrosaurine ceratopsid dinosaur that lived during the Late Cretaceous Period of what is now North America, about 74.2 million years ago. The first fossils of Achelousaurus were collected in Montana in 1987, by a team led by Jack Horner, with more finds made in 1989. In 1994, Achelousaurus horneri was described and named by Scott D. Sampson; the generic name means "Achelous lizard", in reference to the Greek deity Achelous, and the specific name refers to Horner. The genus is known from a few specimens consisting mainly of skull material from individuals, ranging from juveniles to adults.
Centrosaurus is a genus of centrosaurine ceratopsian dinosaur from Campanian age of Late Cretaceous Canada. Their remains have been found in the Dinosaur Park Formation, dating from 76.5 to 75.5 million years ago.
Arrhinoceratops is a genus of herbivorous ceratopsian dinosaur. The name was coined as its original describer concluded it was special because the nose-horn was not a separate bone, however further analysis revealed this was based on a misunderstanding. It lived during the latest Campanian/earliest Maastrichtian stage of the Late Cretaceous, predating its famous relative Triceratops by a few million years, although it was contemporary with Anchiceratops. Its remains have been found in Canada.
Monoclonius is an extinct genus of herbivorous ceratopsian dinosaur found in the Late Cretaceous layers of the Judith River Formation in Montana, United States, and the uppermost rock layers of the Dinosaur Park Formation in Alberta, Canada dated to between 75 and 74.6 million years ago.
Avaceratops is a genus of small herbivorous ceratopsian dinosaurs which lived during the late Campanian in what are now the Northwest United States. Most fossils come from the Judith River Formation.
Ceratops is a dubious genus of herbivorous ceratopsian dinosaur which lived during the Late Cretaceous. Its fossils have been found in the Judith River Formation in Montana. Although poorly known, Ceratops is important in the history of dinosaurs, since it is the type genus for which both the Ceratopsia and the Ceratopsidae have been named.
Medusaceratops is an extinct genus of centrosaurine ceratopsian dinosaur known from the Late Cretaceous Judith River Formation of Montana, northern United States. It contains a single species, Medusaceratops lokii.
Centrosaurinae is a subfamily of ceratopsid, a group of large quadrupedal ornithischian dinosaur. Centrosaurine fossil remains are known primarily from the northern region of Laramidia but isolated taxa have been found in China and Utah as well.
Sinoceratops is an extinct genus of ceratopsian dinosaur that lived from 73.5 to 66 million years ago during the latter part of the Cretaceous Period in what is now Shandong province in China. It was named in 2010 by Xu Xing et al. for three skulls from Zhucheng, China. The name of its type species Sinoceratops zhuchengensis means "Chinese horned face from Zhucheng", after the location of its discovery.
Kosmoceratops is a genus of ceratopsid dinosaur that lived in North America about 76–75.9 million years ago during the Late Cretaceous period. Specimens were discovered in Utah in the Kaiparowits Formation of the Grand Staircase–Escalante National Monument in 2006 and 2007, including an adult skull and postcranial skeleton and partial subadults. In 2010, the adult was made the holotype of the new genus and species Kosmoceratops richardsoni; the generic name means "ornate horned face", and the specific name honors Scott Richardson, who found the specimens. The find was part of a spate of ceratopsian discoveries in the early 21st century, and Kosmoceratops was considered significant due to its elaborate skull ornamentation.
Nasutoceratops is genus of ceratopsid dinosaur that lived in North America during the Late Cretaceous period, about 76.0–75.5 million years ago. The first known specimens were discovered in Utah in the Kaiparowits Formation of the Grand Staircase–Escalante National Monument (GSENM) from 2006 onwards, including a subadult skull with a partial postcranial skeleton and rare skin impressions and two other partial skulls. In 2013, the subadult was made the holotype of the new genus and species Nasutoceratops titusi; the generic name means "large-nosed horned face", and the specific name honors the paleontologist Alan L. Titus for his work at the GSENM. The dinosaur was noted for its large nose in news reports, and later featured in Jurassic World films.
Coronosaurus is a genus of centrosaurine ceratopsian dinosaurs which lived in the Late Cretaceous, in the middle Campanian stage. Its remains, two bone beds, were discovered by Phillip J. Currie in the Oldman Formation of Alberta, Canada, and its type and only species, Coronosaurus brinkmani, was first described in 2005, as a new species within the genus Centrosaurus. Later studies questioned the presence of a direct relationship, and in 2012 it was named as a separate genus. Coronosaurus means "crowned lizard", coming from "corona", Latin for crown, and "sauros", Greek for lizard; this name refers to the unique, crown-like shape of the horns on the top of its frill.
This timeline of ceratopsian research is a chronological listing of events in the history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bony frills, and, later, spectacular horns. The first scientifically documented ceratopsian fossils were described by Edward Drinker Cope starting in the 1870s; however, the remains were poorly preserved and their true nature was not recognized. Over the next several decades, Cope named several such genera and species. Cope's hated rival, Othniel Charles Marsh, also described ceratopsian remains. In 1887, Marsh mistook a Triceratops horn for one belonging to a new species of prehistoric Bison. Marsh also named the eponymous genus Ceratops in 1888. The next year, he named the most famous ceratopsian, Triceratops horridus. It was the discovery of Triceratops that illuminated the ceratopsian body plan, and he formally named the Ceratopsia in 1890.
Stellasaurus is a genus of centrosaurine ceratopsid dinosaur that lived in Montana during the Late Cretaceous. The type and only species is Stellasaurus ancellae. Its remains have been found in the Late Campanian age Two Medicine Formation, the same geological unit which its relatives Rubeosaurus, Einiosaurus, and Achelousaurus were discovered in.