Turanoceratops Temporal range: Turonian, | |
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Life restoration of Turanoceratops tardabilis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Neornithischia |
Clade: | † Ceratopsia |
Clade: | † Coronosauria |
Superfamily: | † Ceratopsoidea |
Genus: | † Turanoceratops Nesov et al., 1989 |
Species: | †T. tardabilis |
Binomial name | |
†Turanoceratops tardabilis Nesov et al., 1989 | |
Turanoceratops ("Turan horned face") is a genus of herbivorous ceratopsian dinosaur from the late Cretaceous Bissekty Formation of Uzbekistan. The fossils dated from the mid-late Turonian stage, roughly 90 million years ago. The skull bore a pair of long brow horns like those seen in the Ceratopsidae, although Turanoceratops appears to have been transitional between earlier ceratopsians and ceratopsids, and not a ceratopsid itself.
From the 1920s onwards, Soviet scientists discovered fragmentary fossils near Dzharakuduk in the district Navoi Viloyat, leading them to the conclusion that some ceratopsid must have been present. In 1988, paleontologist Lev Aleksandrovich Nesov based on these published the name Turanoceratops tardabilis, [1] but did not provide a description so that for the time being it remained a nomen nudum . In 1989, Nesov, L.F. Kaznysjkina and Gennady Olegovich Cherepanov validly named the type species Turanoceratops tardabilis. The generic name is a combination of Turan, an old Persian name for Turkestan, the general region of the finds, and ~ceratops, "horned face", a usual suffix in ceratopian names. The specific name means "retarding" in Latin, referring to the protracted research. [2]
The holotype, CCMGE No. 251/12457, consists of a damaged left maxilla, the tooth-bearing upper jaw bone. Other fossils have been referred but some of these later were proven to have belonged to other types of dinosaur. A braincase (specimen CCGME 628/12457) was shown to be of a sauropod. The material includes postorbitals with brow horn cores, a maxilla, teeth, a predentary and limb elements.
In 2004, Peter Dodson considered it a nomen dubium , [3] but in 2009 Hans-Dieter Sues and Alexander Averianov concluded that it was a valid taxon. [4]
Turanoceratops was a relatively small animal. In 2010, Gregory S. Paul estimated its length at two metres, its weight at 175 kilogrammes. [5]
Sues and Averianov in 2009 determined some typically basal traits, indicating a lower position in the evolutionary tree, and some more derived traits, showing a higher position. Basal traits are the variably developed secondary ridges on the tooth crowns and the possession of just two or three teeth per tooth position. Derived traits, relative to the more basal Zuniceratops , are the increase of the number of teeth to two or three and the possession of two roots per teeth. More general derived traits are the exclusion of the frontal bone from the upper rim of the eye socket and the presence of incipient cavities in the skull roof. [4]
Turanoceratops belonged to the Ceratopsia (the name is Greek for "horned face"), a group of herbivorous dinosaurs with parrot-like beaks which thrived in North America and Asia during the Cretaceous Period, which ended 66 million years ago. All ceratopsians became extinct at the end of this period. A 2009 study led by Hans-Dieter Sues analysed additional fossil material of Turanoceratops and concluded that, contrary to expectations, it represented a true (though "transitional") member of the family Ceratopsidae. If correct, it would represent an Asian ceratopsid. At the time of publication this would have been unique, as all other ceratopsids known to that point were from North America. [4] Some scientists, such as Andrew Farke, disagreed with Sues' findings. Farke and colleagues ran an independent phylogenetic analysis of the new Turanoceratops fossils and found that it was a close relative of Ceratopsidae (the immediate sister group) but was not a true member of that clade. [6] Sues and Alexander Averianov criticised that analysis, arguing that Farke and colleagues misinterpreted or mis-coded some characteristics of the fossil in their analysis. [7] However, Xu et al. (2010) conducted a phylogenetic analysis and concluded that Turanoceratops was more derived than Zuniceratops and was outside of Ceratopsidae, because it lacks several important synapomorphies of this group.
Turanoceratops, like all ceratopsians, was a herbivore. During the Cretaceous, flowering plants were "geographically limited on the landscape"[ clarification needed ][ citation needed ], and so it is likely that this dinosaur fed on the predominant plants of the era: ferns and conifers. It would have used its sharp ceratopsian beak to bite off the leaves or needles.[ citation needed ]
Triceratops is a genus of chasmosaurine ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous period, about 68 to 66 million years ago in what is now western North America. It was one of the last-known non-avian dinosaurs and lived until the Cretaceous–Paleogene extinction event 66 million years ago. The name Triceratops, which means 'three-horned face', is derived from the Greek words trí- meaning 'three', kéras meaning 'horn', and ṓps meaning 'face'.
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7 million years ago. The last ceratopsian species, Triceratops prorsus, became extinct during the Cretaceous–Paleogene extinction event, 66 million years ago.
Ceratopsidae is a family of ceratopsian dinosaurs including Triceratops, Centrosaurus, and Styracosaurus. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are known from western North America, which formed the island continent of Laramidia during most of the Late Cretaceous. Ceratopsids are characterized by beaks, rows of shearing teeth in the back of the jaw, elaborate nasal horns, and a thin parietal-squamosal shelf that extends back and up into a frill. The group is divided into two subfamilies—Chasmosaurinae and Centrosaurinae. The chasmosaurines are generally characterized by long, triangular frills and well-developed brow horns. The centrosaurines had well-developed nasal horns or nasal bosses, shorter and more rectangular frills, and elaborate spines on the back of the frill.
Marginocephalia is a clade of ornithischian dinosaurs that is characterized by a bony shelf or margin at the back of the skull. These fringes were likely used for display. This clade was officially defined in the PhyloCode by Daniel Madzia and colleagues in 2021 as "the smallest clade containing Ceratops montanus, Pachycephalosaurus wyomingensis, and Triceratops horridus". There are two clades included in Marginocephalia: the thick-skulled Pachycephalosauria and the horned Ceratopsia. All members of Marginocephalia were primarily herbivores. They basally used gastroliths to aid in digestion of tough plant matter until they convergently evolved tooth batteries in Neoceratopsia and Pachycephalosauria. Marginocephalia first evolved in the Jurassic Period and became more common in the Cretaceous. They are basally small facultative quadrupeds while derived members of the group are large obligate quadrupeds. Primitive marginocephalians are found in Asia, but the group migrated upwards into North America.
Pentaceratops is a genus of herbivorous ceratopsid dinosaur from the late Cretaceous Period of what is now North America. Fossils of this animal were first discovered in 1921, but the genus was named in 1923 when its type species, Pentaceratops sternbergii, was described. Pentaceratops lived around 76–73 million years ago, its remains having been mostly found in the Kirtland Formation in the San Juan Basin in New Mexico. About a dozen skulls and skeletons have been uncovered, so anatomical understanding of Pentaceratops is fairly complete. One exceptionally large specimen later became its own genus, Titanoceratops, due to its more derived morphology, similarities to Triceratops, and lack of unique characteristics shared with Pentaceratops.
Nedoceratops is a controversial genus of ceratopsid dinosaur from the Late Cretaceous period Lance Formation of North America. It is known only from a single skull discovered in Wyoming. Its status is the subject of ongoing debate among paleontologists: some authors consider Nedoceratops a valid, distinct taxon, while others consider it to be an unusual specimen of Triceratops.
Styracosaurus is an extinct genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of North America. It had four to six long parietal spikes extending from its neck frill, a smaller jugal horn on each of its cheeks, and a single horn protruding from its nose, which may have been up to 60 centimeters long and 15 centimeters wide. The function or functions of the horns and frills have been debated for many years.
Achelousaurus is a genus of centrosaurine ceratopsid dinosaur that lived during the Late Cretaceous Period of what is now North America, about 74.2 million years ago. The first fossils of Achelousaurus were collected in Montana in 1987, by a team led by Jack Horner, with more finds made in 1989. In 1994, Achelousaurus horneri was described and named by Scott D. Sampson; the generic name means "Achelous lizard", in reference to the Greek deity Achelous, and the specific name refers to Horner. The genus is known from a few specimens consisting mainly of skull material from individuals, ranging from juveniles to adults.
Zuniceratops is a genus of ceratopsian dinosaurs that lived during the Turonian stage of the Late Cretaceous in what is now New Mexico, United States. Only a single species is known, Zuniceratops christopheri.
Leptoceratops is a genus of ceratopsian dinosaur from the Late Cretaceous of North America. First found in Alberta in 1910, the type species Leptoceratops gracilis was named in 1914 by Barnum Brown for a partial skull and skeleton of two individuals found in the Scollard Formation of Alberta. Additional specimens found in the Scollard include one complete and two mostly complete skeletons together, uncovered in 1947 by Charles M. Sternberg. Specimens from Montana that were among the earliest referred to Leptoceratops have since been moved to their own genera Montanoceratops and Cerasinops, while new specimens of L. gracilis include bonebed remains from the Hell Creek Formation of Montana and a partial skeleton from the Lance Formation of Wyoming. Together with related taxa, Leptoceratops is the eponymous genus of the family Leptoceratopsidae. Leptoceratops is known from more than ten individuals, all from Maastrichtian deposits of Alberta, Montana and Wyoming, representing the entire skeleton.
Anchiceratops is an extinct genus of chasmosaurine ceratopsid dinosaur that lived approximately 72 to 71 million years ago during the latter part of the Cretaceous Period in what is now Alberta, Canada. Anchiceratops was a medium-sized, heavily built, ground-dwelling, quadrupedal herbivore that could grow up to an estimated 4.3 metres (14 ft) long. Its skull featured two long brow horns and a short horn on the nose. The skull frill was elongated and rectangular, its edges adorned by coarse triangular projections. About a dozen skulls of the genus have been found.
Arrhinoceratops is a genus of herbivorous ceratopsian dinosaur. The name was coined as its original describer concluded it was special because the nose-horn was not a separate bone, however further analysis revealed this was based on a misunderstanding. It lived during the latest Campanian/earliest Maastrichtian stage of the Late Cretaceous, predating its famous relative Triceratops by a few million years, although it was contemporary with Anchiceratops. Its remains have been found in Canada.
Asiaceratops is a genus of herbivorous ceratopsian dinosaur. It lived during the Early-Late Cretaceous. The type species, A. salsopaludalis is known from Uzbekistan, while A. sulcidens is known from China and Mongolia.
Asiamericana is a dubious genus of coelurosaur known only from isolated teeth found in the Bissekty Formation of Uzbekhistan. It was named to recognize the occurrence of similar fossil teeth in Central Asia and North America. These regions once formed a connected land mass, during the Cretaceous period.
Kulceratops is a genus of ceratopsian dinosaur from the Early Cretaceous. It lived in the late Albian stage. It is one of the few ceratopsians known from this period. However, the fossils from this genus have been sparse: only jaw and tooth fragments have been found so far.
The Bissekty Formation is a geologic formation and Lagerstätte which crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period. Laid down in the mid to late Turonian, it is dated to about 92 to 90 Ma.
Ojoceratops is a genus of ceratopsian dinosaur which lived in what is now New Mexico, United States. Ojoceratops fossils have been recovered from strata of the Ojo Alamo Formation, dating to the late Cretaceous period. The type species is Ojoceratops fowleri.
Vagaceratops is a genus of herbivorous ceratopsian dinosaur. It is a chasmosaurine ceratopsian which lived during the Late Cretaceous period in what is now Alberta. Its fossils have been recovered from the Upper Dinosaur Park Formation. It is sometimes included in the genus Chasmosaurus as Chasmosaurus irvinensis instead of being recognized as its own genus.
This timeline of ceratopsian research is a chronological listing of events in the history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, bony frills, and, later, spectacular horns. The first scientifically documented ceratopsian fossils were described by Edward Drinker Cope starting in the 1870s; however, the remains were poorly preserved and their true nature was not recognized. Over the next several decades, Cope named several such genera and species. Cope's hated rival, Othniel Charles Marsh, also described ceratopsian remains. In 1887, Marsh mistook a Triceratops horn for one belonging to a new species of prehistoric Bison. Marsh also named the eponymous genus Ceratops in 1888. The next year, he named the most famous ceratopsian, Triceratops horridus. It was the discovery of Triceratops that illuminated the ceratopsian body plan, and he formally named the Ceratopsia in 1890.