Ctenosauriscidae

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Ctenosauriscids
Temporal range: Early-Middle Triassic, 247.5–237  Ma
Ctenosauriscus koeneni holotype.png
Vertebrae of the ctenosauriscid Ctenosauriscus koeneni
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauria
Clade: Pseudosuchia
Superfamily: Poposauroidea
Family: Ctenosauriscidae
Kuhn, 1964
Genera

Ctenosauriscidae is an extinct family of pseudosuchian archosaurs within the clade Poposauroidea. [1] Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest Olenekian to Anisian stages). [1] All species had large "sails" on their backs. Ctenosauriscids are among some of the earliest archosaurs and represent the first global radiation of the group. [2]

Genera

GenusStatusAgeLocationUnitNotesImages

Arizonasaurus

Valid

Anisian

Flag of the United States.svg  USA

Moenkopi Formation

Arizonasaurus Arizonasaurus BW.jpg
Arizonasaurus
Ctenosauriscus Ctenosauriscus BW.jpg
Ctenosauriscus
Some fossils of the "Waldhaus ctenosauriscid" Ctenosauriscus koeneni.JPG
Some fossils of the "Waldhaus ctenosauriscid"

Bromsgroveia

Valid

Anisian

Flag of the United Kingdom.svg  United Kingdom

Bromsgrove Sandstone Formation

Bystrowisuchus

Valid

Olenekian

Flag of Russia.svg  Russia

Lipovskaya Formation

Ctenosauriscus

Valid

Olenekian

Flag of Germany.svg  Germany

Solling Formation

Ctenosauriscus is one of the oldest archosaurs known to date. [2]

Hypselorhachis

Valid

Anisian

Flag of Tanzania.svg  Tanzania

Manda Formation

"Waldhaus" ctenosauriscid

Valid

Anisian

Flag of Germany.svg  Germany

Röt Formation

Xilousuchus

Valid

Olenekian

Flag of the People's Republic of China.svg  China

Heshanggou Formation

Xilousuchus is one of the oldest archosaurs known to date. [2]

Phylogeny

Ctenosauriscidae was named by Oskar Kuhn in 1964 to include the genus Ctenosauriscus . [3] It is a stem-based taxon defined by Richard J. Butler, Stephen L. Brusatte, Mike Reich, Sterling J. Nesbitt, Rainer R. Schoch and Jahn J. Hornung in 2011 as "the most inclusive clade containing Ctenosauriscus koeneni but not Poposaurus gracilis , Effigia okeeffeae , Postosuchus kirkpatricki , Crocodylus niloticus , Ornithosuchus longidens , or Aetosaurus ferratus ". [2] The cladograms below follows a 2011 analysis by Butler et al., the first based on Brusatte et al. 2010 [4] and the second based on Nesbitt 2011. [1]

Arizonasaurus model, Museum am Lowentor, Stuttgart Ctenosauriscid model.jpg
Arizonasaurus model, Museum am Lowentor, Stuttgart
Poposauroidea
Poposauroidea

Related Research Articles

<span class="mw-page-title-main">Archosaur</span> Group of diapsids broadly classified as reptiles

Archosauria is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term, which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives; and Avemetatarsalia, which includes birds and their extinct relatives.

<span class="mw-page-title-main">Archosauriformes</span> Clade of reptiles

Archosauriformes is a clade of diapsid reptiles encompassing archosaurs and some of their close relatives. It was defined by Jacques Gauthier (1994) as the clade stemming from the last common ancestor of Proterosuchidae and Archosauria. Phil Senter (2005) defined it as the most exclusive clade containing Proterosuchus and Archosauria. Archosauriforms are a branch of archosauromorphs which originated in the Late Permian and persist to the present day as the two surviving archosaur groups: crocodilians and birds.

<span class="mw-page-title-main">Rauisuchia</span> Informal group of Triassic archosaurs with pillar-erect posture

"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.

<i>Arizonasaurus</i> Extinct genus of reptiles

Arizonasaurus was a ctenosauriscid archosaur from the Middle Triassic. Arizonasaurus is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, Arizonasaurus babbitti, was named by Samuel Paul Welles in 1947.

<i>Riojasuchus</i> Extinct genus of reptiles

Riojasuchus is an extinct genus of ornithosuchid archosaur from the Late Triassic (Norian) of Argentina. Ornithosuchidae was a widespread family of facultatively bipedal pseudosuchians with adaptations for scavenging. Riojasuchus is notable as one of the youngest and most complete members of the family. The type and only known species, Riojasuchus tenuisceps, was named and described by José Bonaparte in 1967. It was one of the first of many well-preserved Triassic archosaurs to be discovered in Argentina. The holotype specimen, PVL 3827, was found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina.

<span class="mw-page-title-main">Pseudosuchia</span> Clade of reptiles

Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.

<i>Ticinosuchus</i> Extinct species of reptile

Ticinosuchus is an extinct genus of suchian archosaur from the Middle Triassic of Switzerland and Italy.

<i>Lotosaurus</i> Extinct genus of reptiles

Lotosaurus is an extinct genus of sail-backed poposauroid known from Hunan Province of central China.

<i>Bromsgroveia</i> Extinct genus of reptiles

Bromsgroveia is an extinct genus of predatory ctenosauriscid archosaur from the Middle Triassic Bromsgrove Sandstone of England. Ctenosauriscids were a group of rauisuchians that was related to the ancestors of modern crocodiles and alligators.

<i>Batrachotomus</i> Genus of reptiles

Batrachotomus is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian stage of the Middle Triassic period, around 242 to 237 million years ago. Batrachotomus was described by palaeontologist David J. Gower 22 years after its discovery.

<i>Ctenosauriscus</i> Extinct genus of reptiles

Ctenosauriscus is an extinct genus of sail-backed poposauroid archosaur from Early Triassic deposits of Lower Saxony in northern Germany. It gives its name to the family Ctenosauriscidae, which includes other sail-backed poposauroids such as Arizonasaurus. Fossils have been found in latest Olenekian deposits around 247.5-247.2 million years old, making it one of the first known archosaurs.

Yarasuchus is an extinct genus of avemetatarsalian archosaur that lived during the Anisian stage of the Middle Triassic of India. The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the Middle Triassic Yerrapalli Formation. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other. The type and only species is Y. deccanensis. Yarasuchus was a quadruped roughly 2–2.5 metres (6.6–8.2 ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of Yarasuchus were slender, and more like those of ornithodirans.

<i>Fasolasuchus</i> Extinct genus of reptiles

Fasolasuchus is an extinct genus of loricatan. Fossils have been found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina that date back to the Norian stage of the Late Triassic, making it one of the last "rauisuchians" to have existed before the order became extinct at the end of the Triassic.

<span class="mw-page-title-main">Suchia</span> Clade of reptiles

Suchia is a clade of archosaurs containing the majority of pseudosuchians. It was defined as the least inclusive clade containing Aetosaurus ferratus, Rauisuchus tiradentes, Prestosuchus chiniquensis, and Crocodylus niloticus by Nesbitt (2011). Generally the only pseudosuchian group which is omitted from Suchia is the family Ornithosuchidae, although at least one analysis classifies ornithosuchids as close relatives of erpetosuchids and aetosaurs. Phytosaurs are also excluded from Suchia, although it is not certain whether they qualify as pseudosuchians in the first place.

<span class="mw-page-title-main">Loricata</span> Extinct clade of reptiles

Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as Postosuchus and Prestosuchus. More specifically, Loricata includes Crocodylomorpha and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs.

Vytshegdosuchus is an extinct genus of paracrocodylomorph archosaur known from the Early Triassic Yarenskian Gorizont of the Komi Republic of the European section of Russia. It contains a single species, Vytshegdosuchus zheshartensis. Vytshegdosuchus was named by Andreii Sennikov in 1988.

<span class="mw-page-title-main">Poposauroidea</span> Extinct clade of reptiles

Poposauroidea is a clade of advanced pseudosuchians. It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as Qianosuchus and Lotosaurus. It excludes most large predatory quadrupedal "rauisuchians" such as rauisuchids and "prestosuchids". Those reptiles are now allied with crocodylomorphs in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes Poposaurus and its close relatives.

Xilousuchus is an extinct genus of poposauroid from lower Triassic deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is Xilousuchus sapingensis. Wu (1981) referred Xilousuchus to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt et al. (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found Xilousuchus to be a poposauroid which is most closely related to Arizonasaurus. Xilousuchus is the oldest archosaur to date, although Ctenosauriscus and Vytshegdosuchus might be even older by less than one million year. Since Xilousuchus is a suchian archosaur, its early age suggests that most of the major groups of archosaurs developed by the Early Triassic, soon after the appearance of the first archosaur.

<span class="mw-page-title-main">Röt Formation</span>

The Röt Formation or Rötton Formation, or Upper Buntsandstein, is a geologic formation of the Buntsandstein in Germany. It preserves fossils dating back to the Middle Triassic Epoch. The formation overlies the Plattenstein and Solling Formations and is overlain by the Jena Formation.

Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.

References

  1. 1 2 3 Sterling J. Nesbitt (2011). "The early evolution of archosaurs: relationships and the origin of major clades". Bulletin of the American Museum of Natural History. 352: 1–292. doi: 10.1206/352.1 . hdl:2246/6112. S2CID   83493714.
  2. 1 2 3 4 Richard J. Butler; Stephen L. Brusatte; Mike Reich; Sterling J. Nesbitt; Rainer R. Schoch; Jahn J. Hornung (2011). "The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation". PLOS ONE. 6 (10): e25693. Bibcode:2011PLoSO...625693B. doi: 10.1371/journal.pone.0025693 . PMC   3194824 . PMID   22022431.
  3. Oskar Kuhn (1964). "Ungelöste Probleme der Stammesgeschichte der Amphibien und Reptilien" [Unsolved problems of the phylogeny of amphibians and reptiles]. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg. 118–119: 293–325.
  4. Brusatte, S.L.; Benton, M.J.; Desojo, J.B.; Langer, M.C. (2010). "The higher-level phylogeny of Archosauria (Tetrapoda: Diapsida)" (PDF). Journal of Systematic Palaeontology. 8 (1): 3–47. Bibcode:2010JSPal...8....3B. doi:10.1080/14772010903537732. hdl: 20.500.11820/24322ff3-e80e-45f2-8d53-d35fd104195c . S2CID   59148006.