Erpetosuchus

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Erpetosuchus
Temporal range: Late Triassic 237–208.5  Ma
Erpetosuchus fossils.jpg
Fossils described in 1894
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauria
Clade: Pseudosuchia
Clade: Suchia
Family: Erpetosuchidae
Genus: Erpetosuchus
Newton, 1894
Species
  • E. grantiNewton, 1894 (type)

Erpetosuchus is an extinct genus of pseudosuchian from the Late Triassic. The type species of Erpetosuchus is E. granti. It was first described by E. T. Newton in 1894 for remains found in northeastern Scotland, including four specimens from the latest Carnian Lossiemouth Sandstone Formation. [1] Additional remains of Erpetosuchus have been found in the New Haven Formation of Connecticut in the eastern United States, although they were not attributed to the species E. granti. The relationship of Erpetosuchus to other archosaurs is uncertain. In 2000 and 2002, it was considered a close relative of the group Crocodylomorpha, which includes living crocodylians and many extinct relatives. However, this relationship was questioned in a 2012 analysis that found the phylogenetic placement of Erpetosuchus to be very uncertain.

Contents

Discovery

Life restoration Erpetosuchus granti.png
Life restoration

The first remains of Erpetosuchus were found in the Lossiemouth Sandstone Formation in Scotland, dating back to the late Carnian stage of the Late Triassic. The holotype specimen is BMNH R3139 and consists of a skull and a partial postcranial skeleton.

During a field trip in 1995 to the lower part of the New Haven Formation in Connecticut, American palaeontologist Paul E. Olsen discovered a partial skull that, after preparation and description in 2000 (Olsen et al. 2000), was referable to Erpetosuchus. This was the first record of Erpetosuchus outside Scotland. The specimen has been given the number AMNH 29300, and besides the right side of the skull, also has some vertebrae and indeterminate bones associated. Dating of the lower portion of the New Haven Formation indicates a Norian age. [2]

Classification

Restoration of the head Erpetosuchus.jpg
Restoration of the head

Erpetosuchus granti was originally assigned to Thecodontia, but that name is no longer considered valid in scientific literature because it is a paraphyletic group of early archosaurs.

A more recent phylogenetic analysis by Olsen et al. (2000) found E. granti to be the sister-taxon to the Crocodylomorpha. These were united in a clade by the following synapomorphies:

Benton and Walker (2002) found the same sister-group relationship and proposed the name Bathyotica for the clade containing Erpetosuchus and Crocodylomorpha.

Nesbitt and Butler (2012) included Erpetosuchus within a more comprehensive phylogenetic analysis and found it to group with the archosaur Parringtonia from the Middle Triassic of Tanzania. Both were part of the clade Erpetosuchidae. Nesbitt and Butler did not find support for the sister-group relationship between Erpetosuchus and Crocodylomorpha. Instead, erpetosuchids formed a polytomy or unresolved evolutionary relationship at the base of Archosauria along with several other groups. It could take many positions within Archosauria, but none were as a sister taxon of Crocodylomorpha. [3]

Related Research Articles

<span class="mw-page-title-main">Rauisuchia</span> Informal group of Triassic archosaurs with pillar-erect posture

"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.

Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution

<span class="mw-page-title-main">Aetosaur</span> Extinct order of heavily armoured reptiles

Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.

<i>Gracilisuchus</i> Genus of fossil reptiles

Gracilisuchus is an extinct genus of tiny pseudosuchian from the Late Triassic of Argentina. It contains a single species, G. stipanicicorum, which is placed in the clade Suchia, close to the ancestry of crocodylomorphs. Both the genus and the species were first described by Alfred Romer in 1972.

<span class="mw-page-title-main">Pseudosuchia</span> Clade of reptiles

Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.

<span class="mw-page-title-main">Prestosuchidae</span> Extinct family of reptiles

Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.

<span class="mw-page-title-main">Sphenosuchia</span> Suborder of reptiles

Sphenosuchia is a suborder of basal crocodylomorphs that first appeared in the Triassic and occurred into the Middle Jurassic. Most were small, gracile animals with an erect limb posture. They are now thought to be ancestral to crocodyliforms, a group which includes all living crocodilians.

<i>Luperosuchus</i> Extinct genus of reptiles

Luperosuchus is an extinct genus of loricatan pseudosuchian reptile which contains only a single species, Luperosuchus fractus. It is known from the Chañares Formation of Argentina, within strata belonging to the latest Ladinian stage of the late Middle Triassic, or the earliest Carnian of the Late Triassic. Luperosuchus was one of the largest carnivores of the Chañares Formation, although its remains are fragmentary and primarily represented by a skull with similarities to Prestosuchus and Saurosuchus.

Yarasuchus is an extinct genus of avemetatarsalian archosaur that lived during the Anisian stage of the Middle Triassic of India. The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the Middle Triassic Yerrapalli Formation. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other. The type and only species is Y. deccanensis. Yarasuchus was a quadruped roughly 2–2.5 metres (6.6–8.2 ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of Yarasuchus were slender, and more like those of ornithodirans.

Euscolosuchus is an extinct genus of suchian archosaurs from the Late Triassic of Virginia. It is probably an aetosauriform, as the sister taxon to Acaenasuchus and a relative of aetosaurs.

<i>Parringtonia</i> Extinct genus of reptiles

Parringtonia is an extinct genus of Triassic archosaur within the family Erpetosuchidae, known from the type species Parringtonia gracilis. It is known from a single specimen, NHMUK R8646, found from the Anisian-age Manda Formation of Tanzania. This specimen, like most archosaur material from the Manda Formation, is fragmentary, including only a maxilla and a few postcranial bones. They show similarities with those of another archosaur called Erpetosuchus, known from the Middle Triassic of Scotland and the eastern United States. The phylogenetic placement of Parringtonia and Erpetosuchus are uncertain; some studies placed them close to the group Crocodylomorpha, which includes all modern crocodylians and many extinct forms that diversified after the Triassic, but this relationship has more recently been questioned.

<span class="mw-page-title-main">Suchia</span> Clade of reptiles

Suchia is a clade of archosaurs containing the majority of pseudosuchians. It was defined as the least inclusive clade containing Aetosaurus ferratus, Rauisuchus tiradentes, Prestosuchus chiniquensis, and Crocodylus niloticus by Nesbitt (2011). Generally the only pseudosuchian group which is omitted from Suchia is the family Ornithosuchidae, although at least one analysis classifies ornithosuchids as close relatives of erpetosuchids and aetosaurs. Phytosaurs are also excluded from Suchia, although it is not certain whether they qualify as pseudosuchians in the first place.

<span class="mw-page-title-main">Loricata</span> Extinct clade of reptiles

Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as Postosuchus and Prestosuchus. More specifically, Loricata includes Crocodylomorpha and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs.

<span class="mw-page-title-main">Doswelliidae</span> Extinct family of reptiles

Doswelliidae is an extinct family of carnivorous archosauriform reptiles that lived in North America and Europe during the Middle to Late Triassic period. Long represented solely by the heavily-armored reptile Doswellia, the family's composition has expanded since 2011, although two supposed South American doswelliids were later redescribed as erpetosuchids. Doswelliids were not true archosaurs, but they were close relatives and some studies have considered them among the most derived non-archosaurian archosauriforms. They may have also been related to the Proterochampsidae, a South American family of crocodile-like archosauriforms.

<span class="mw-page-title-main">Bathyotica</span> Clade of reptiles

Bathyotica is a clade of crurotarsan archosaurs that includes the superorder Crocodylomorpha and its sister taxon Erpetosuchus, a small Triassic suchian. Bathyotica was named in a 2002 phylogenetic study of Erpetosuchus. The genus was found to be closely related to crocodylomorphs, and Bathyotica was erected to encompass both taxa.

<span class="mw-page-title-main">Paracrocodylomorpha</span> Clade of reptiles

Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing Poposaurus and Crocodylus niloticus. Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic.

<span class="mw-page-title-main">Proterochampsia</span> Extinct clade of reptiles

Proterochampsia is a clade of early archosauriform reptiles from the Triassic period. It includes the Proterochampsidae and probably also the Doswelliidae. Nesbitt (2011) defines Proterochampsia as a stem-based taxon that includes Proterochampsa and all forms more closely related to it than Euparkeria, Erythrosuchus, Passer domesticus, or Crocodylus niloticus. Therefore, the inclusion of Doswelliidae in it is dependent upon whether Doswellia and Proterochampsa form a monophyletic group to the exclusion of Archosauria and other related groups.

<span class="mw-page-title-main">Erpetosuchidae</span> Extinct family of reptiles

Erpetosuchidae is an extinct family of pseudosuchian archosaurs. Erpetosuchidae was named by D. M. S. Watson in 1917 to include Erpetosuchus. It includes the type species Erpetosuchus granti from the Late Triassic of Scotland, Erpetosuchus sp. from the Late Triassic of eastern United States and Parringtonia gracilis from the middle Middle Triassic of Tanzania; the group might also include Dyoplax arenaceus from the Late Triassic of Germany, Archeopelta arborensis and Pagosvenator candelariensis from Brazil and Tarjadia ruthae from Argentina.

<i>Asperoris</i> Extinct genus of reptiles

Asperoris is an extinct genus of archosauriform reptile known from the Middle Triassic Manda Beds of southwestern Tanzania. It is the first archosauriform known from the Manda Beds that is not an archosaur. However, its relationships with other non-archosaurian archosauriforms are uncertain. It was first named by Sterling J. Nesbitt, Richard J. Butler and David J. Gower in 2013 and the type species is Asperoris mnyama. Asperoris means "rough face" in Latin, referring to the distinctive rough texture of its skull bones.

<span class="mw-page-title-main">Aetosauriformes</span> Extinct clade of reptiles

Aetosauriformes is an extinct clade of early-diverging pseudosuchians. It includes the aetosaurs, a group of heavily armoured and at least partially herbivorous pseudosuchians, as well as the closely related genera Acaenasuchus, Euscolosuchus and Revueltosaurus.

References

  1. "Palaeos Mesozoic: Triassic: Late Triassic: Carnian - 3". Archived from the original on 2013-06-17. Retrieved 2012-05-22.
  2. Olsen, P. E.; Sues, H. D.; Norell, M. A. (2000). "First record ofErpetosuchus(Reptilia: Archosauria) from the Late Triassic of North America". Journal of Vertebrate Paleontology. 20 (4): 633. doi:10.1671/0272-4634(2000)020[0633:FROERA]2.0.CO;2. S2CID   8229773.
  3. Nesbitt, S. J.; Butler, R. J. (2012). "Redescription of the archosaur Parringtonia gracilis from the Middle Triassic Manda beds of Tanzania, and the antiquity of Erpetosuchidae". Geological Magazine. 150 (2): 225–238. doi:10.1017/S0016756812000362. S2CID   232175772.