Vjushkovisaurus Temporal range: Middle Triassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Clade: | Archosauriformes |
Genus: | † Vjushkovisaurus Ochev, 1982 |
Type species | |
†Vjushkovisaurus berdjanensis Ochev, 1982 |
Vjushkovisaurus is an extinct genus of Middle Triassic archosauriform. It is known from the Anisian-aged Donguz Gorizont in Sol-Iletsk, Orenburg Oblast, Russia. The genus was named in 1982, with the type species being V. berdjanensis. [1] Material has been collected in the Berdyanka II locality from a fossil assemblage called the Eryosuchus Fauna along the Berdyanka River, specifically in a sand-carbonate concretion in the upper part of the main river channel. [2] Vjushkovisaurus is known only from the holotype PIN 2865/62 (formerly SGU 104/3871), a partial postcranial skeleton which consists of 12 presacral vertebrae, left humerus, ribs, a fragment of the coracoid and a fragment of the fibula. [3]
Most rauisuchids have a crest on the ilium called the supra-acetabular process that overlies the head of the femur, allowing them to have a "pillar-erect" stance. [4] [5] As an early rauisuchid Vjushkovisaurus lacks this crest, but it does have a small thickening on the surface of the ilium. [6] The humerus differs from other better known rauisuchids in that it is shorter and broader. However, it has been suggested that the only known humerus of Vjushkovisaurus may actually belong to an aetosaur. [7]
More recently, Nesbitt (2009) argued that Vjushkovisaurus represent most probably a valid non-archosaurian archosauriform. According to his analysis, the original diagnosis of Vjushkovisaurus by Ochev (1982) consisted of some of the plesiomorphies for Archosauriformes and characters with wide distributions throughout archosauriforms. For example, elongated anterior cervical vertebrae are present in the poposauroids, Yarasuchus , and in the non-archosaurian archosauriforms Proterosuchus and Guchengosuchus . Other features which are typically thought to be associated with skeletal pneumaticity, are also present in Guchengosuchus, Erythrosuchus , and in several archosaurs. Additionally, the shape and placement of the deltopectoral crests of the humerus of Vjushkovisaurus are typical of basal archosauriforms. The ectepicondylar groove on the humerus reported by Ochev is present in Prolacerta , Proterosuchus, phytosaurs, aetosaurs and several paracrocodylomorphs. Gower and Sennikov (2000) reported that the posterior cervical vertebrae bear well-defined facets for three-headed ribs. According to Nesbitt (2009), this trait is absent in Archosauria with the possible exception of Arizonasaurus and Poposaurus , but was used as a synapomorphy of Erythrosuchia in Parrish (1992) phylogenetic analysis. [3]
Other vertebrates from the Berdyanka II locality that lived alongside Vjushkovisaurus include capitosauroid temnospondyls and the therapsid Nothogomphodon , one of the last therocephalians. [2] Vjushkovisaurus was likely a top predator in its environment along other large carnivores including Dongusuchus and Erythrosuchus . [8]
Archosauriformes is a clade of diapsid reptiles encompassing archosaurs and some of their close relatives. It was defined by Jacques Gauthier (1994) as the clade stemming from the last common ancestor of Proterosuchidae and Archosauria. Phil Senter (2005) defined it as the most exclusive clade containing Proterosuchus and Archosauria. Gauthier as part of the Phylonyms (2020) defined the clade as the last common ancestor and all descendants of Gallus, Alligator, and Proterosuchus. Archosauriforms are a branch of archosauromorphs which originated in the Late Permian and persist to the present day as the two surviving archosaur groups: crocodilians and birds.
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.
Rauisuchidae is a group of large predatory Triassic archosaurs. Some disagreement exists over which genera should be included in the Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid families. Rauisuchidae in the modern sense was defined by Sterling Nesbitt in 2011 as the most inclusive clade containing Rauisuchus tiradentes, but not Prestosuchus chiniquensis, Poposaurus gracilis, or Crocodylus niloticus. In this modern sense, rauisuchids are recovered as members of the clade Loricata, being the sister taxon of Crocodylomorpha, and being more derived than taxa such as Prestosuchus and Batrachotomus. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as Scythosuchus and Tsylmosuchus are considered to be within the family.
Erythrosuchidae are a family of large basal archosauriform carnivores that lived from the later Early Triassic (Olenekian) to the early Middle Triassic (Anisian).
Proterosuchidae is an early family of basal archosauriforms whose fossils are known from the Late Permian and the Early Triassic. The highest diversity of genera is known from European Russia, but fossils are also known from South Africa, India, China, Australia, Brazil and possibly Argentina. The name comes from Greek πρότερο- ("first") and σοῦχος ("crocodile").
Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.
Teleocrater is a genus of avemetatarsalian archosaur from the Middle Triassic Manda Formation of Tanzania. The name was coined by English paleontologist Alan Charig in his 1956 doctoral dissertation, but was only formally published in 2017 by Sterling Nesbitt and colleagues. The genus contains the type and only species T. rhadinus. Uncertainty over the affinities of Teleocrater have persisted since Charig's initial publication; they were not resolved until Nesbitt et al. performed a phylogenetic analysis. They found that Teleocrater is most closely related to the similarly enigmatic Yarasuchus, Dongusuchus, and Spondylosoma in a group that was named the Aphanosauria. Aphanosauria was found to be the sister group of the Ornithodira, the group containing dinosaurs and pterosaurs.
Garjainia is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.5–2 metres (4.9–6.6 ft) long. It contained two species, Garjainia prima from the Yarengian/Yarkenskian Supergorizont of Russia, and Garjainia madiba from the Burgersdorp Formation of South Africa. "Vjuskovia triplicostata", a name assigned to some erythrosuchid fossils from Russia, has been synonymized with Garjainia prima.
Turfanosuchus is a genus of archosauriform reptile, likely a gracilisuchid archosaur, which lived during the Middle Triassic (Anisian) of northwestern China. The type species, T. dabanensis, was described by C.C. Young in 1973, based on a partially complete but disarticulated fossil skeleton found in the Kelamayi Formation of the Turfan Basin.
Archosaurus is an extinct genus of carnivorous proterosuchid archosauriform reptile. Its fossils are dated to the latest Permian of Russia and Poland, it is one of the earliest known archosauriforms. The type and only species is Archosaurus rossicus, known from several fragmentary specimens which cumulatively represent parts of the skull and cervical vertebrae. It would have been 3 metres (9.8 ft) long when fully grown.
Yarasuchus is an extinct genus of avemetatarsalian archosaur that lived during the Anisian stage of the Middle Triassic of India. The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the Middle Triassic Yerrapalli Formation. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other. The type and only species is Y. deccanensis. Yarasuchus was a quadruped roughly 2–2.5 metres (6.6–8.2 ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of Yarasuchus were slender, and more like those of ornithodirans.
Dongusuchus is an extinct genus of archosaur. Fossils have been found from the Donguz Formation outcropping on the banks of the Donguz River in the Orenburg Oblast of Russia. They are associated with a fossil assemblage called the Eryosuchus Fauna, named after the capitosaurid Eryosuchus, the most common organism found from the assemblage. The locality dates back to the Anisian and early Ladinian stages of the Middle Triassic.
Mandasuchus is an extinct genus of loricatan pseudosuchian from the Manda Formation of Tanzania, which dates back to the Anisian stage of the Middle Triassic. Although this genus was first mentioned by Alan Charig in 1956, a formal description was not published until 2018.
Stagonosuchus is an extinct genus of paracrocodylomorph, possibly a loricatan synonymous with Prestosuchus or a poposauroid. Fossils have been found from the Late Triassic Manda Formation in Tanzania that are Anisian in age.
Tsylmosuchus is an extinct genus of proterosuchid archosauriform reptile known from Western Russia. Fossils referred to Tsylmosuchus occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs (pseudosuchians) such as Mandasuchus. As a result, Tsylmosuchus was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so Tsylmosuchus has more recently been interpreted as an indeterminate archosauriform. Although three species of Tsylmosuchus have been named, they lack diagnostic traits and are probably not distinct from each other. In 2023, Tsylmosuchus was reinterpreted as a proterosuchid, specifically a member of the subfamily Chasmatosuchinae.
Youngosuchus is an extinct genus of archosaur from the Middle Triassic of China. The type species is Y. sinensis. Y. sinensis was first described in 1973 as a new species of the erythrosuchid Vjushkovia. In 1985, it was reassigned as its own genus of rauisuchid. A 1992 study supported the original classification of Youngosuchus sinensis as an erythrosuchid, but more recent studies classify it as a "rauisuchian"-grade loricatan archosaur completely unrelated to Vjushkovia, which is most likely a synonym of Garjainia.
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing Poposaurus and Crocodylus niloticus. Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic.
Vonhuenia is an extinct genus of proterosuchid, a basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.
Wangisuchus is an extinct genus of archosauriform reptile from the Middle Triassic of China that is known from fragmentary fossil jaw bones. These bones were found at the Hsishihwa locality in the upper Ermaying Formation, which dates to the late Anisian stage about 242 million years ago. Wangisuchus was named in 1964 by Chinese paleontologist Yang Zhongjian, who described a single species, Wangisuchus tzeyii, on the basis of these bones.
Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.