| Ticinosuchus Temporal range: Middle Triassic, | |
|---|---|
 | |
| Holotype of Ticinosuchus ferox | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Clade: | Archosauria |
| Clade: | Pseudosuchia |
| Clade: | Suchia |
| Genus: | † Ticinosuchus Krebs, 1965 |
| Species: | †T. ferox |
| Binomial name | |
| †Ticinosuchus ferox Krebs, 1965 | |
Ticinosuchus is an extinct genus of suchian archosaur from the Middle Triassic (Anisian - Ladinian) of Switzerland and Italy. [1]
One of only a handful of fossil reptiles that have been found in Switzerland, Ticinosuchus (meaning "Ticino crocodile" due to its origin from the Swiss canton Ticino) was about 3 metres (10 ft) long, and its whole body, even the belly, was covered in thick, armoured scutes. These scutes were sometimes considered to have been staggered, alternating between several rows. [2] However, some studies refute this claim, instead purporting that the scutes were aligned in neat rows, with a one-to-one assignment of scutes to vertebrae. [3] The structure of the hips shows that its legs were placed under the body almost vertically. Coupled with the development of a calcaneus and a specialized ankle joint, this would have made Ticinosuchus a fast runner, unlike most earlier reptiles. [4] Ticinosuchus is thought to be very close to or possible even the same species that made the Cheirotherium trace fossils found in Germany. It too shows a narrow track-way, similar to that of Ticinosuchus. It is one of the most famous fossils of Besano. [5]
Fish scales have been preserved in the abdomen of the specimen. This was likely indicative of a piscivorous diet. [6] Ticinosuchus shares many similarities with paracrocodylomorphs, such as certain adaptations of the ischium [1] and possibly (but not certainly) hyposphene-hypantrum articulations. [7] [8]
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.
Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution
Euparkeria is an extinct genus of archosauriform reptile from the Triassic of South Africa. Euparkeria is close to the ancestry of Archosauria, the reptile group that includes crocodilians, pterosaurs, and dinosaurs.
Scleromochlus is an extinct genus of small pterosauromorph archosaurs from the Late Triassic Lossiemouth Sandstone of Scotland. The genus contains the type and only species Scleromochlus taylori, named by Arthur Smith Woodward in 1907.
Riojasuchus is an extinct genus of ornithosuchid archosaur from the Late Triassic (Norian) of Argentina. Ornithosuchidae was a widespread family of facultatively bipedal pseudosuchians with adaptations for scavenging. Riojasuchus is notable as one of the youngest and most complete members of the family. The type and only known species, Riojasuchus tenuisceps, was named and described by José Bonaparte in 1967. It was one of the first of many well-preserved Triassic archosaurs to be discovered in Argentina. The holotype specimen, PVL 3827, was found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina.
Lewisuchus is a genus of archosaur that lived during the Late Triassic. As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs. Lewisuchus was about 1 metre (3.3 ft) long. Fossils have been found in the Chañares Formation of Argentina. It exhibited osteoderms along its back.
Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age, and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid Staurikosaurus, the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.
Yarasuchus is an extinct genus of avemetatarsalian archosaur that lived during the Anisian stage of the Middle Triassic of India. The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the Middle Triassic Yerrapalli Formation. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other. The type and only species is Y. deccanensis. Yarasuchus was a quadruped roughly 2–2.5 metres (6.6–8.2 ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of Yarasuchus were slender, and more like those of ornithodirans.
Sterling Nesbitt is an American paleontologist best known for his work on the origin and early evolutionary patterns of archosaurs. He is currently an associate professor at Virginia Tech in the Department of Geosciences.
Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids comprised an early diverging clade or a paraphyletic grade within ornithischian dinosaurs. Silesaurids have a consistent general body plan, with a fairly long neck and legs and possibly quadrupedal habits, but most silesaurids are heavily fragmentary nonetheless. Furthermore, they occupied a variety of ecological niches, with early silesaurids being carnivorous and later taxa having adaptations for specialized herbivory. As indicated by the contents of referred coprolites, Silesaurus may have been insectivorous, feeding selectively on small beetles and other arthropods.
Stagonosuchus is an extinct genus of loricatan, or possibly a species of Prestosuchus. Fossils have been found from the Late Triassic Manda Formation in Tanzania that are Anisian in age.
Suchia is a clade of archosaurs containing the majority of pseudosuchians. It was defined as the least inclusive clade containing Aetosaurus ferratus, Rauisuchus tiradentes, Prestosuchus chiniquensis, and Crocodylus niloticus by Nesbitt (2011). Generally the only pseudosuchian group which is omitted from Suchia is the family Ornithosuchidae, although at least one analysis classifies ornithosuchids as close relatives of erpetosuchids and aetosaurs. Phytosaurs are also excluded from Suchia, although it is not certain whether they qualify as pseudosuchians in the first place.
Doswelliidae is an extinct family of carnivorous archosauriform reptiles that lived in North America and Europe during the Middle to Late Triassic period. Long represented solely by the heavily-armored reptile Doswellia, the family's composition has expanded since 2011, although two supposed South American doswelliids were later redescribed as erpetosuchids. Doswelliids were not true archosaurs, but they were close relatives and some studies have considered them among the most derived non-archosaurian archosauriforms. They may have also been related to the Proterochampsidae, a South American family of crocodile-like archosauriforms.
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing Poposaurus and Crocodylus niloticus. Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic.
The Manda Formation is a Middle Triassic (Anisian?) or possibly Late Triassic (Carnian?) geologic formation in Tanzania. It preserves fossils of many terrestrial vertebrates from the Triassic, including some of the earliest dinosauromorph archosaurs. The formation is often considered to be Anisian in age according to general tetrapod biochronology hypotheses and correlations to the Cynognathus Assemblage Zone of South Africa. However, some recent studies cast doubt to this age, suggesting that parts deposits may actually be younger (Carnian) in age.
The Ermaying Formation is a geological formation of Anisian age in north-central China. It is found across much of the Ordos Basin, at outcrops within the provinces of Shaanxi, Shanxi, and Inner Mongolia. It is composed of up to 600 m thick sequence of mudstone and sandstone, overlying the Heshanggou Formation and underlying the Tongchuan Formation. In the southern part of the Ordos Basin, the Zhifang Formation is equivalent to the Ermaying Formation.
Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.
The Besano Formation is a geological formation in the southern Alps of northwestern Italy and southern Switzerland. This formation, a short but fossiliferous succession of dolomite and black shale, is famous for its preservation of Middle Triassic (Anisian–Ladinian) marine life including fish and aquatic reptiles. It is exposed in the vicinity of Monte San Giorgio and is among the formations responsible for the area being designated as a UNESCO World Heritage Site. In Switzerland, it is also known as the Grenzbitumenzone. The Anisian-Ladinian boundary lies in the upper part of the Besano Formation.
