Vivaron

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Vivaron
Temporal range: Late Triassic, Norian
~212  Ma
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Vivaron.jpg
Right maxilla of the holotype
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Family: Rauisuchidae
Genus: Vivaron
Lessner et al., 2016
Species:
V. haydeni
Binomial name
Vivaron haydeni
Lessner et al, 2016

Vivaron is a genus of rauisuchid known from the Late Triassic (middle Norian) Chinle Formation in New Mexico. It is the second rauisuchid known from the southwestern United States, and it highlights the wide biogeographic range similar rauisuchid taxa occupied during the Late Triassic across Pangaea, despite the varied faunal assemblages at different latitudes. [1]

Contents

Discovery

Stratigraphic and geographic location of the Hayden Quarry Hayden Quarry.png
Stratigraphic and geographic location of the Hayden Quarry

Vivaron was named in 2016 from material collected at the Hayden Quarry at Ghost Ranch, New Mexico in 2009. [2] The locality is part of the Chinle Formation in the Petrified Forest Member, and dates to the middle Norian ~212 Ma, possibly representing one of the youngest known rauisuchids. Prior to its description, all Late Triassic rauisuchid material from Texas, Arizona and New Mexico had been referred to Postosuchus kirkpatricki . However, the rauisuchid remains from Hayden Quarry could be clearly distinguished from Postosuchus, and was erected as a new taxon Vivaron haydeni.

The generic epithet was named for Vivaron, a mythical 30 foot rattlesnake spirit that was said to live under Mesa Huerfano (Orphan Mesa) at Ghost Ranch, [3] and the specific name in honour of John Hayden, a hiker who discovered the Hayden Quarry in 2002 where the material for Vivaron was collected. [1]

Description

Vivaron is known only from three jaw bones, associated skull fragments and referred hip bones. Comparisons with related taxa allow for an estimated length of between 12 and 18 feet (3.7 and 5.5 m), similar in size and appearance to other rauisuchids. [2]

Vivaron can be distinguished from other rauisuchids by the presence of two prongs on the posterior portion of the maxilla where it articulates with the jugal. It is also unique among rauisuchids by possessing five alveoli in the premaxilla, similar to the state in early crocodylomorphs but in contrast with other rauisuchids, which only have four alveoli (however, a reconstruction of a Postosuchus skull (UCMP A269) figured in Long & Murry 1995 [4] (Fig. 121) possibly shows a fifth alveolus in the left premaxilla). Vivaron was noted to share a number of characteristics with Teratosaurus , to which it is believed to be closely related to, particularly features of the maxilla and antorbital fossa. The referred hip elements also share similarities to the referred hips of Teratosaurus, potentially supporting their referral to each taxon, despite neither being directly associated with the holotype. [1]

Classification

Referred premaxilla Premaxilla of Vivaron.png
Referred premaxilla

Vivaron was found to be a member of Rauisuchidae by Lessner et al. 2016 in their phylogenetic analysis, with no changes to the relationships of pseudosuchians found in a previous analysis (Nesbitt, 2011). [5] The analysis recovered Rauisuchidae as monophyletic but completely unresolved, with all six species in a polytomy. Surveys of the interrelationships within Rauisuchidae found P. kirkpatricki as the sister taxon of Polonosuchus , and Rauisuchus as the sister taxon to the rest of Rauisuchidae.

The presence of Vivaron in the southwestern United States in addition to Postosuchus has implications for the classification of numerous other rauisuchid material from these locations, as almost all of it had been referred to the single species Postosuchus kirkpatricki based on their geographic location. Vivaron demonstrates the need for more detailed morphological analysis of all rauisuchid remains in the southwestern United States when determining their assignment within the group, and not refer them to a taxon based only on geographic distribution.

The inferred close relationship between Vivaron and Teratosaurus also has implications for the biogeography of middle Norian Pangaea. During the late Carnian to early Norian, very similar faunal associations were present across Pangaea, all of which included rauisuchid taxa. However, by the middle Norian faunal assemblages were becoming disparate, particularly evident between high-latitude locations in Europe (including sauropodomorphs, phytosaurs, aetosaurs and turtles) and the lower-latitude southern United States (including dinosauromorphs, metoposaurs, phytosaurs, aetosaurs and non-archosaur archosauromorphs). The close relationship between Vivaron and Teratosaurus demonstrates that despite being widely geographically separated and belonging to distinct faunal assemblages, there was a clear biogeographic link between the Chinle Formation and high-latitude Pangaea, and that rauisuchids were able to maintain a widespread distribution into the middle Norian. [1]

Related Research Articles

<span class="mw-page-title-main">Rauisuchia</span> Informal group of Triassic archosaurs with pillar-erect posture

"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.

Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution. The name "phytosaur" means "plant reptile", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters.

<i>Postosuchus</i> Genus of reptiles

Postosuchus, meaning "Crocodile from Post", is an extinct genus of rauisuchid reptiles comprising two species, P. kirkpatricki and P. alisonae, that lived in what is now North America during the Late Triassic. Postosuchus is a member of the clade Pseudosuchia, the lineage of archosaurs that includes modern crocodilians. Its name refers to Post Quarry, a place in Texas where many fossils of the type species, P. kirkpatricki, were found. It was one of the apex predators of its area during the Triassic, larger than the small dinosaur predators of its time. It was a hunter which probably preyed on large bulky herbivores like dicynodonts and many other creatures smaller than itself.

<span class="mw-page-title-main">Aetosaur</span> Extinct order of heavily armoured reptiles

Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.

<i>Revueltosaurus</i> Extinct genus of reptiles

Revueltosaurus is an extinct genus of suchian pseudosuchian from Late Triassic deposits of New Mexico, Arizona and North Carolina, United States. Many specimens, mostly teeth, have been assigned to Revueltosaurus over the years. Currently, three species are included in this genus, all of which were originally thought to represent monospecific genera of basal ornithischian dinosaurs. It was 1 meter long.

<i>Shuvosaurus</i> Genus of beaked reptile

Shuvosaurus is a genus of beaked reptile from the Late Triassic of western Texas. Despite looking superficially similar to a theropod dinosaur, it is actually more closely related to crocodilians.

Spinosuchus is an extinct genus of trilophosaurid allokotosaur from the Late Triassic of Texas, southern United States. It has been assigned to a variety of groups over its history, from coelophysid dinosaur to pseudosuchian to uncertain theropod dinosaur and to Proterosuchidae. This uncertainty is not unusual, given that it was only known from a poorly preserved, wall-mounted, partial vertebral column of an animal that lived in a time of diverse, poorly known reptile groups. However, newly collected material and recent phylogenetic studies of early archosauromorphs suggest that it represents an advanced trilophosaurid very closely related to Trilophosaurus.

<span class="mw-page-title-main">Poposauridae</span> Extinct family of reptiles

Poposauridae is a family of large carnivorous archosaurs which lived alongside dinosaurs during the Late Triassic. They were around 2.5 to 5 metres long. Poposaurids are known from fossil remains from North and South America. While originally believed to be theropod dinosaurs, cladistic analysis has shown them to be more closely related to crocodiles.

<i>Gracilisuchus</i> Genus of fossil reptiles

Gracilisuchus is an extinct genus of tiny pseudosuchian from the Late Triassic of Argentina. It contains a single species, G. stipanicicorum, which is placed in the clade Suchia, close to the ancestry of crocodylomorphs. Both the genus and the species were first described by Alfred Romer in 1972.

<span class="mw-page-title-main">Pseudosuchia</span> Clade of reptiles

Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.

<span class="mw-page-title-main">Rauisuchidae</span> Extinct family of reptiles

Rauisuchidae is a group of large predatory Triassic archosaurs. Some disagreement exists over which genera should be included in the Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid families. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as Scythosuchus and Tsylmosuchus are considered to be within the family.

<span class="mw-page-title-main">Prestosuchidae</span> Extinct family of reptiles

Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.

<i>Dromomeron</i> Extinct genus of reptiles

Dromomeron is a genus of lagerpetid avemetatarsalian which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from Late Triassic-age rocks of the Southwestern United States and northwestern Argentina. It is described as most closely related to the earlier Lagerpeton of Argentina, but was found among remains of true dinosaurs like Chindesaurus, indicating that the first dinosaurs did not immediately replace related groups.

Redondasaurus is an extinct genus or subgenus of phytosaur from the Late Triassic of the southwestern United States. It was named by Hunt & Lucas in 1993, and contains two species, R. gregorii and R. bermani. It is the youngest and most evolutionarily-advanced of the phytosaurs.

The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico. It is one of several formations encompassed by the Dockum Group.

Euscolosuchus is an extinct genus of suchian archosaurs from the Late Triassic of Virginia. It is probably an aetosauriform, as the sister taxon to Acaenasuchus and a relative of aetosaurs.

<i>Heptasuchus</i> Extinct genus of reptiles

Heptasuchus is an extinct genus of loricatan pseudosuchian known from the Middle or Late Triassic upper Chugwater Group of Wyoming, United States. It contains a single species, Heptasuchus clarki, the first formally recognized "rauisuchian" or loricatan pseudosuchian from North America.

<span class="mw-page-title-main">Aetosauriformes</span> Extinct clade of reptiles

Aetosauriformes is an extinct clade of early-diverging pseudosuchians. It includes the aetosaurs, a group of heavily armoured and at least partially herbivorous pseudosuchians, as well as the closely related genera Acaenasuchus, Euscolosuchus and Revueltosaurus.

Land vertebrate faunachrons (LVFs) are biochronological units used to correlate and date terrestrial sediments and fossils based on their tetrapod faunas. First formulated on a global scale by Spencer G. Lucas in 1998, LVFs are primarily used within the Triassic Period, though Lucas later designated LVFs for other periods as well. Eight worldwide LVFs are defined for the Triassic. The first two earliest Triassic LVFs, the Lootsbergian and Nonesian, are based on South African synapsids and faunal assemblage zones estimated to correspond to the Early Triassic. These are followed by the Perovkan and Berdyankian, based on temnospondyl amphibians and Russian assemblages estimated to be from the Middle Triassic. The youngest four Triassic LVFs, the Otischalkian, Adamanian, Revueltian, and Apachean, are based on aetosaur and phytosaur reptiles common in the Late Triassic of the southwestern United States.

References

  1. 1 2 3 4 Emily J. Lessner; Michelle R. Stocker; Nathan D. Smith; Alan H. Turner; Randall B. Irmis; Sterling J. Nesbitt (2016). "A new rauisuchid (Archosauria, Pseudosuchia) from the Upper Triassic (Norian) of New Mexico increases the diversity and temporal range of the clade". PeerJ. 4: e2336. doi: 10.7717/peerj.2336 . PMC   5018681 . PMID   27651983.
  2. 1 2 Mackay, Steven (6 September 2016). "Undergraduate researcher leads study naming a new species of reptile from 212 million years ago". Virginia Tech News. Retrieved 7 September 2016.
  3. Poling-Kempes, Lesley (2005). Ghost Ranch. Tucson: University of Arizona Press, 2005. p. 14. ISBN   0816523479.
  4. Robert A. Long; Philip A. Murry (1995). "Late Triassic (Carnian and Norian) Tetrapods from the Southwestern United States". Bulletin of the New Mexico Museum of Natural History and Science. 4: 121.
  5. Nesbitt, Sterling J. (2011). "The Early Evolution of Archosaurs: Relationships and the Origin of Major Clades". Bulletin of the American Museum of Natural History . 352: 1–292. doi: 10.1206/352.1 . hdl:2246/6112. S2CID   83493714.