Protanystropheus Temporal range: Middle Triassic, | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Clade: | † Dinocephalosauridae |
Genus: | † Protanystropheus Sennikov, 2011 |
Type species | |
†Protanystropheus antiquus | |
Synonyms | |
? Thecodontosaurus primusHuene, 1908 |
Protanystropheus is an extinct genus of archosauromorph from the Middle Triassic (Anisian stage) of Poland, Germany, Austria and the Netherlands. It was named by Sennikov in 2011 and the type species is Protanystropheus antiquus, first described in 1908 by German paleontologist Friedrich von Huene under the name Tanystropheus antiquus [1] (some authors still prefer to include this species within Tanystropheus [2] [3] ). Sennikov (2011) referred to Protanystropheus several vertebrae, including those belonging to " Thecodontosaurus " primus, [1] but such a referral has later been questioned, because these specimens may represent other basal archosauromorphs. [2]
Dinocephalosaurus is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, D. orientalis, which was named by Li in 2003. Unlike other long-necked protorosaurs, Dinocephalosaurus convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative Pectodens, which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, Dinocephalosaurus probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as suggested by the poorly-ossified and paddle-like limbs which would have prevented it from going ashore.
Tanystropheus is an extinct genus of archosauromorph reptile which lived during the Triassic Period in Europe, Asia, and North America. It is recognisable by its extremely elongated neck, longer than the torso and tail combined. The neck was composed of 13 vertebrae strengthened by extensive cervical ribs. Tanystropheus is one of the most well-described non-archosauriform archosauromorphs, known from numerous fossils, including nearly complete skeletons. Some species within the genus may have reached a total length of 6 meters (20 ft), making Tanystropheus the longest non-archosauriform archosauromorph as well. Tanystropheus is the namesake of the family Tanystropheidae, a clade collecting many long-necked Triassic archosauromorphs previously described as "protorosaurs" or "prolacertiforms".
Thecodontosaurus is a genus of herbivorous basal sauropodomorph dinosaur that lived during the late Triassic period.
Lepidosauromorpha is a group of reptiles comprising all diapsids closer to lizards than to archosaurs. The only living sub-group is the Lepidosauria, which contains two subdivisions, Squamata, which contains lizards and snakes, and Rhynchocephalia, the only extant species of which is the tuatara.
Proterosuchus is an extinct genus of archosauriform reptiles that lived during the Early Triassic. It contains three valid species: the type species P. fergusi and the referred species P. alexanderi and P. goweri. All three species lived in what is now South Africa. The genus was named in 1903 by the South African paleontologist Robert Broom. The genus Chasmatosaurus is a junior synonym of Proterosuchus.
Macrocnemus is an extinct genus of archosauromorph reptile known from the Middle Triassic of Europe and China. Macrocnemus is a member of the Tanystropheidae family and includes three species. Macrocnemus bassanii, the first species to be named and described, is known from the Besano Formation and adjacent paleontological sites in the Italian and Swiss Alps. Macrocnemus fuyuanensis, on the other hand, is known from the Zhuganpo Formation in southern China. A third species, Macrocnemus obristi, is known from the Prosanto Formation of Switzerland and is characterized by gracile limbs. The name Macrocnemus is Greek for "long tibia".
Tsylmosuchus is an extinct genus of proterosuchid archosauriform reptile known from Western Russia. Fossils referred to Tsylmosuchus occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs (pseudosuchians) such as Mandasuchus. As a result, Tsylmosuchus was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so Tsylmosuchus has more recently been interpreted as an indeterminate archosauriform. Although three species of Tsylmosuchus have been named, they lack diagnostic traits and are probably not distinct from each other. In 2023, Tsylmosuchus was reinterpreted as a proterosuchid, specifically a member of the subfamily Chasmatosuchinae.
Vritramimosaurus is an extinct genus of large early archosauromorph. Although originally placed in the family Prolacertidae, recent studies on archosauromorph relationships doubt the validity of the family, at least in its broadest sense. Fossils have been found from Early Triassic deposits of the Rassypnaya locality in Orenburg Oblast, Russia. Rassypnaya is located on the Obshchy Syrt, a plateau in the European part of Russia that extends southwest of the Urals toward the Volga River. Vritramimosaurus is similar to the later genus Malutinisuchus, also from Rassypnaya but present in Middle Triassic deposits.
Malutinisuchus is an extinct genus of Archosauromorph. The genus was named in 1986 with the description of the type species M. gratus. Malutinisuchus is known from Ladinian-age Middle Triassic deposits in the Bukobay and Rassypnaya localities in Orenburg Oblast, Russia. In Russia, deposits of this age are referred to the Bukobay Gorizont.
Augustaburiania is an extinct genus of tanystropheid archosauromorph from the latest Early Triassic of Volgograd Region, western Russia. All specimens were recovered in the right slope of the Don River valley from the Lipovskaya Formation. It was named by Sennikov in 2011 and the type species is Augustaburiania vatagini. Augustaburiania is the oldest known tanystropheid.
Youngosuchus is an extinct genus of archosaur from the Middle Triassic of China. The type species is Y. sinensis. Y. sinensis was first described in 1973 as a new species of the erythrosuchid Vjushkovia. In 1985, it was reassigned as its own genus of rauisuchid. A 1992 study supported the original classification of Youngosuchus sinensis as an erythrosuchid, but more recent studies classify it as a "rauisuchian"-grade loricatan archosaur completely unrelated to Vjushkovia, which is most likely a synonym of Garjainia.
Protorosauria is an extinct, likely paraphyletic group of basal archosauromorph reptiles from the latest Middle Permian to the end of the Late Triassic of Asia, Europe and North America. It was named by the English anatomist and paleontologist Thomas Henry Huxley in 1871 as an order, originally to solely contain Protorosaurus. Other names which were once considered equivalent to Protorosauria include Prolacertiformes and Prolacertilia.
Coelodontognathus is an extinct genus of reptile from the Early Triassic of European Russia. It was originally described as a procolophonid parareptile in 1967 but was reclassified as a possible trilophosaurid archosauromorph in 2008. The genus includes two species: the type species C. donensis and C. ricovi. C. donensis is known from the holotype PIN 4173/129 and the referred PIN 4173/130, and C. ricovi is known from the holotype PIN 4173/127 and the referred PIN 4173/128, all of which represent dentaries that are housed at the Paleontological Institute, Russian Academy of Sciences. Another dentary, SGU 104/3105, originally referred to C. donensis was reassigned to its own genus and species Vitalia grata by Ivakhnenko, 1973. The fossils have been found at the Donskaya Luka Locality near the village of Sirotinskaya and the Don River in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. Like Coelodontognathus, Vitalia which is known from the same locality was also first identified as a procolophonid and later reclassified as a trilophosaurid. Coelodontognathus and Vitalia are similar to procolophonids in that they have wide teeth but differs from them in that they have tooth roots set deep into the jaws.
Tanystropheidae is an extinct family of mostly marine archosauromorph reptiles that lived throughout the Triassic Period, often considered to be "protorosaurs". They are characterized by their long, stiff necks formed from elongated cervical vertebrae with very long cervical ribs. Some tanystropheids such as Tanystropheus had necks that were several meters long, longer than the rest of their bodies.
Eorasaurus is an extinct genus of archosauromorph reptile known from the middle Late Permian of Tatarstan, European Russia. It contains a single species, Eorasaurus olsoni. When originally described by Sennikov (1997), Eorasaurus was identified as an early archosauromorph and assigned to the family Protorosauridae, Ezcurra et al. (2014) and Ezcurra (2016) later reclassified Eorasaurus and placed it within the group Archosauriformes. Eorasaurus is based solely on scant fossil material from the neck region, and is thus considered an unstable taxon in phylogenetic analyses. If Eorasaurus is an archosauriform, it would be the oldest known member of the group and would pre-date the previous record holder.
Doniceps is an extinct genus of reptile from the Early Triassic of European Russia known from the type species D. lipovensis. It was solely known from the holotype premaxilla 104/3106 housed at Saratov University, however the specimen is probably lost. It was collected at the Donskaya Luka Locality near the village of Sirotinskaya in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. The generic name refers to the nearby Don River added the Greek suffix for "head", -ceps. The specific name is derived from the name of the type locality Donskaya Luka, also known as Lipovaya Balka. Named by Otschev and Rykov in 1968 as an archosauromorph, Arkhangelskii & Sennikov (2008) classified the taxon as a possible trilophosaurid. Doniceps is thought to be similar to Coelodontognathus and Vitalia, both of which are known exclusively from the same locality and were first identified as procolophonids but later reclassified as trilophosaurids.
Vitalia is an extinct genus of reptile from the Early Triassic of European Russia known from the type species V. grata. It is known from the holotype dentary PIN 4173/126 as well as two additional dentaries PIN 1043/627 and 1043/628, all housed at the Paleontological Institute, Russian Academy of Sciences. The type dentary was originally included in the hypodigm of Coelodontognathus donensis named by the notable Russian vertebrate paleontologist Vitaliy Georgiyevich Ochev in 1967. Ivakhnenko (1973) separated the specimen and gave it its own genus and species name in light of the new material, which he named in honor of Ochev. The dentaries of Vitalia were collected at the Donskaya Luka Locality near the village of Sirotinskaya in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. Like Coelodontognathus, Vitalia was originally described as a procolophonid parareptile in 1973, but Arkhangelskii & Sennikov (2008) reclassified the taxon as a possible trilophosaurid archosauromorph. Vitalia is thought to be similar to the possible trilophosaurids Coelodontognathus and Doniceps, both of which are known exclusively from the same locality. Coelodontognathus and Vitalia are similar to procolophonids in that they have wide teeth but differs from them in that they have tooth roots set deep into the jaws.
Ozimek is a genus of sharovipterygid archosauromorph reptile, known from Late Triassic deposits in Poland and closely related to the Kyrgyzstani Sharovipteryx. It contains one species, O. volans, named in 2016 by Jerzy Dzik and Tomasz Sulej. Like Sharovipteryx, Ozimek had long, slender limbs with the hindlimbs longer than the forelimbs; the hindlimbs likely supported gliding membranes as fossilized in Sharovipteryx. Another unusual characteristic was the shoulder girdle, where the massive coracoids formed a shield-like structure covering the bottom of the shoulder region that would have limited mobility. In other respects, such as its long neck, it was a typical member of the non-natural grouping Protorosauria. Phylogenetic analysis has indicated that it, possibly along with Sharovipteryx, may have been an unusual member of the protorosaur group Tanystropheidae, although further study of its anatomy is needed to resolve its precise relationships.
Pectodens is an extinct genus of archosauromorph reptile which lived during the Middle Triassic in China. The type and only species of the genus is P. zhenyuensis, named by Chun Li and colleagues in 2017. It was a member of the Archosauromorpha, specifically part of the unnatural grouping Protorosauria. However, an unusual combination of traits similar and dissimilar to other protorosaurs initially led to confusion over its evolutionary relationships. In 2021, it was placed in a newly-established group, Dinocephalosauridae, along with its closest relative Dinocephalosaurus.
Dinocephalosauridae is an extinct clade of marine and terrestrial archosauromorph reptiles that lived throughout the Triassic period. Like tanystropheids, they are characterized by their long necks, lengthened by either addition of cervical vertebrae or elongation of the individual bones.