Samsarasuchus is known from several vertebrae recovered from the Panchet Formation of West Bengal, India. The formation has been dated to the Induan, the earliest stage of the Triassic period. It was classified in the Chasmatosuchinae, a subfamily of proterosuchids that contains a majority of genera in the family, including the PermianArchosaurus. It is the only valid proterosuchid from India; Ankistrodon, another proterosuchid previously described from the Panchet Formation, is considered a nomen dubium due to its non-diagnostic remains.[1]
The genus name Samsarasuchus references saṃsāra, the eternal cycle of death and rebirth in Indian religions, referring to the recovery of global ecosystems after the Permian-Triassic extinction event. The species name honors paleontologist Pamela Lamplugh Robinson.[1]
The Triassic is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.
Archosauriformes is a clade of diapsid reptiles encompassing archosaurs and some of their close relatives. It was defined by Jacques Gauthier (1994) as the clade stemming from the last common ancestor of Proterosuchidae and Archosauria. Phil Senter (2005) defined it as the most exclusive clade containing Proterosuchus and Archosauria. Archosauriforms are a branch of archosauromorphs which originated in the Late Permian and persist to the present day as the two surviving archosaur groups: crocodilians and birds.
Lystrosaurus is an extinct genus of herbivorous dicynodont therapsids from the late Permian and Early Triassic epochs. It lived in what is now Antarctica, India, China, Mongolia, European Russia and South Africa. Four to six species are currently recognized, although from the 1930s to 1970s the number of species was thought to be much higher. They ranged in size from that of a small dog to 8 feet long.
Ankistrodon is an extinct genus of archosauriform known from the Early Triassic Panchet Formation of India. First thought to be a theropod dinosaur, it was later determined to be a proterosuchid. The type species is A. indicus, described by prolific British zoologist Thomas Henry Huxley in 1865. One authority in the 1970s classified Ankistrodon as a senior synonym of Proterosuchus. Ezcurra (2023) found Ankistrodon to be a nomen dubium, as the teeth are indistinguishable from those of Proterosuchus. A second Indian proterosuchid from the same formation, Samsarasuchus, was also described in the same study, making it the only known valid proterosuchid from India.
Chasmatosuchus was an archosauriform reptile from the early Triassic period of European Russia. One of the earliest described archosauriforms, it was over 2 m long and is thought to have behaved like a modern crocodile. Its mouth had two distinct features: the top of its jaw hooked downwards to aid in holding prey, and the upper palate was lined with a row of teeth—a primitive feature lost in later archosaurs.
Proterosuchidae is an early family of basal archosauriforms whose fossils are known from the Late Permian and the Early Triassic. The highest diversity of genera is known from European Russia, but fossils are also known from South Africa, India, China, Australia, Brazil and possibly Argentina. The name comes from Greek πρότερο- ("first") and σοῦχος ("crocodile").
Proterosuchus is an extinct genus of archosauriform reptiles that lived during the Early Triassic. It contains three valid species: the type species P. fergusi and the referred species P. alexanderi and P. goweri. All three species lived in what is now South Africa. The genus was named in 1903 by the South African paleontologist Robert Broom. The genus Chasmatosaurus is a junior synonym of Proterosuchus.
Archosaurus is an extinct genus of carnivorous proterosuchid archosauriform reptile. Its fossils are dated to the latest Permian of Russia and Poland, it is one of the earliest known archosauriforms. The type and only species is Archosaurus rossicus, known from several fragmentary specimens which cumulatively represent parts of the skull and cervical vertebrae. It would have been 3 metres (9.8 ft) long when fully grown.
Indolyrocephalus is an extinct genus of prehistoric amphibian belonging to the family Trematosauridae. It contains a single species, I. huxleyi, from the Early Triassic Panchet Formation of India. It was initially classified in Gonioglyptus, then into its own genus Indolyrocephalus, and then back into Gonioglyptus, but is presently placed in Indolyrocephalus once again.
Gonioglyptus is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. It is known from the Early Triassic Panchet Formation of India. It contains two species: G. longirostris and G. fragilis. The species G. kokeni from Pakistan has since been reclassified into Aphaneramma.
Pachygonia is an extinct genus of temnospondyl amphibian from the Early Triassic Panchet Formation of India. It contains a single species, P. incurvata. It may potentially be a member of the family Brachyopidae.
Jaikosuchus is an extinct genus of proterosuchid archosauriform. It contains a single species, J. magnus. Fossils have been found from European Russia that date back to the upper Olenekian stage of the Early Triassic.
Tsylmosuchus is an extinct genus of proterosuchid archosauriform reptile known from Western Russia. Fossils referred to Tsylmosuchus occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs (pseudosuchians) such as Mandasuchus. As a result, Tsylmosuchus was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so Tsylmosuchus has more recently been interpreted as an indeterminate archosauriform. Although three species of Tsylmosuchus have been named, they lack diagnostic traits and are probably not distinct from each other. In 2023, Tsylmosuchus was reinterpreted as a proterosuchid, specifically a member of the subfamily Chasmatosuchinae.
Vonhuenia is an extinct genus of proterosuchid, a basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.
Gamosaurus is an extinct genus of proterosuchid archosauriform. It contains a single species, Gamosaurus lozovskii, named by Vitalii Georgievich Ochev in 1979.
Eorasaurus is an extinct genus of archosauromorph reptile known from the middle Late Permian of Tatarstan, European Russia. It contains a single species, Eorasaurus olsoni. When originally described by Sennikov (1997), Eorasaurus was identified as an early archosauromorph and assigned to the family Protorosauridae, Ezcurra et al. (2014) and Ezcurra (2016) later reclassified Eorasaurus and placed it within the group Archosauriformes. Eorasaurus is based solely on scant fossil material from the neck region, and is thus considered an unstable taxon in phylogenetic analyses. If Eorasaurus is an archosauriform, it would be the oldest known member of the group and would pre-date the previous record holder.
Teyujagua is an extinct genus of small, probably semi-aquatic archosauromorph reptile that lived in Brazil during the Early Triassic period. The genus contains the type and only known species, T. paradoxa. It is known from a well-preserved skull, and probably resembled a crocodile in appearance. It was an intermediary between the primitive archosauromorphs and the more advanced Archosauriformes, revealing the mosaic evolution of how the key features of the archosauriform skull were acquired. Teyujagua also provides additional support for a two-phase model of archosauriform radiation, with an initial diversification in the Permian followed by a second adaptive radiation in the Early Triassic.
Pamela Lamplugh Robinson was a British paleontologist who worked extensively on the fauna of the Triassic and Early Jurassic of Gloucestershire and later worked in India on the Mesozoic and Gondwanan fauna. She helped establish the geology unit at the Indian Statistical Institute and directed research in vertebrate paleontology of India in the 1960s.
Manubrantlia was a genus of lapillopsid temnospondyls from the Early Triassic Panchet Formation of India. This genus is only known from a single holotype left jaw, given the designation ISI A 57. Despite the paucity of remains, the jaw is still identifiable as belonging to a relative of Lapillopsis. For example, all three of its coronoid bones possessed teeth, the articular bone is partially visible in lateral (outer) view, and its postsplenial does not contact the posterior meckelian foramen. However, the jaw also possesses certain unique features which justify the erection of a new genus separate from Lapillopsis. For example, the mandible is twice the size of any jaws referred to other lapillopsids. The most notable unique feature is an enlarged "pump-handle" shaped arcadian process at the back of the jaw. This structure is responsible for the generic name of this genus, as "Manubrantlia" translates from Latin to the English expression "pump-handle". The type and only known species of this genus is Manubrantlia khaki. The specific name refers to the greenish-brown mudstones of the Panchet Formation, with a color that had been described as "khaki" by the first British geologists who studied the formation.
The Panchet Formation is an Early Triassic geological formation from the Damodar Valley of India.
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