Vritramimosaurus Temporal range: Early Triassic | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Genus: | † Vritramimosaurus Sennikov, 2005 |
Species | |
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Vritramimosaurus is an extinct genus of large early archosauromorph. Although originally placed in the family Prolacertidae, [1] recent studies on archosauromorph relationships doubt the validity of the family, at least in its broadest sense. Fossils have been found from Early Triassic deposits of the Rassypnaya locality in Orenburg Oblast, Russia. Rassypnaya is located on the Obshchy Syrt, a plateau in the European part of Russia that extends southwest of the Urals toward the Volga River. Vritramimosaurus is similar to the later genus Malutinisuchus , also from Rassypnaya but present in Middle Triassic deposits. [2]
Garjainia is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.5–2 metres (4.9–6.6 ft) long. It contained two species, Garjainia prima from the Yarengian/Yarkenskian Supergorizont of Russia, and Garjainia madiba from the Burgersdorp Formation of South Africa. "Vjuskovia triplicostata", a name assigned to some erythrosuchid fossils from Russia, has been synonymized with Garjainia prima.
Archosaurus is an extinct genus of carnivorous proterosuchid archosauriform reptile. Its fossils are dated to the latest Permian of Russia and Poland, it is one of the earliest known archosauriforms. The type and only species is Archosaurus rossicus, known from several fragmentary specimens which cumulatively represent parts of the skull and cervical vertebrae. It would have been 3 metres (9.8 ft) long when fully grown.
Sarmatosuchus is an extinct genus of archosauriform reptile found in sediments of Middle Triassic age and known from the single species Sarmatosuchus otschevi. It is one of the earliest stem-archosaur species known. The holotype and only specimen was found in lithified river deposits of the Donguz Formation exposed near the Berdyanka River, Orenburg region, Russia. The genus was originally included in the extinct family Proterosuchidae. However, later analysis by David Gower and Andrei Sennikov, the describing author, has removed it from the Proterosuchidae and designated it a basal archosauriform.
Dongusuchus is an extinct genus of archosaur. Fossils have been found from the Donguz Formation outcropping on the banks of the Donguz River in the Orenburg Oblast of Russia. They are associated with a fossil assemblage called the Eryosuchus Fauna, named after the capitosaurid Eryosuchus, the most common organism found from the assemblage. The locality dates back to the Anisian and early Ladinian stages of the Middle Triassic.
Tsylmosuchus is an extinct genus of proterosuchid archosauriform reptile known from Western Russia. Fossils referred to Tsylmosuchus occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs (pseudosuchians) such as Mandasuchus. As a result, Tsylmosuchus was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so Tsylmosuchus has more recently been interpreted as an indeterminate archosauriform. Although three species of Tsylmosuchus have been named, they lack diagnostic traits and are probably not distinct from each other. In 2023, Tsylmosuchus was reinterpreted as a proterosuchid, specifically a member of the subfamily Chasmatosuchinae.
Vjushkovisaurus is an extinct genus of Middle Triassic archosauriform. It is known from the Anisian-aged Donguz Gorizont in Sol-Iletsk, Orenburg Oblast, Russia. The genus was named in 1982, with the type species being V. berdjanensis. Material has been collected in the Berdyanka II locality from a fossil assemblage called the Eryosuchus Fauna along the Berdyanka River, specifically in a sand-carbonate concretion in the upper part of the main river channel. Vjushkovisaurus is known only from the holotype PIN 2865/62, a partial postcranial skeleton which consists of 12 presacral vertebrae, left humerus, ribs, a fragment of the coracoid and a fragment of the fibula.
Malutinisuchus is an extinct genus of Archosauromorph. The genus was named in 1986 with the description of the type species M. gratus. Malutinisuchus is known from Ladinian-age Middle Triassic deposits in the Bukobay and Rassypnaya localities in Orenburg Oblast, Russia. In Russia, deposits of this age are referred to the Bukobay Gorizont.
Vladlenosaurus is an extinct genus of capitosaurian temnospondyl from Russia. It lived during the late Vetlugian. Based on the type of deposits it was found in, Vladlenosaurus probably inhabited lacustrine, or lake, habitats. The type species is V. alexeyevi, named in 2000.
Protanystropheus is an extinct genus of archosauromorph from the Middle Triassic of Poland, Germany, Austria and the Netherlands. It was named by Sennikov in 2011 and the type species is Protanystropheus antiquus, first described in 1908 by German paleontologist Friedrich von Huene under the name Tanystropheus antiquus. Sennikov (2011) referred to Protanystropheus several vertebrae, including those belonging to "Thecodontosaurus" primus, but such a referral has later been questioned, because these specimens may represent other basal archosauromorphs.
Augustaburiania is an extinct genus of tanystropheid archosauromorph from the latest Early Triassic of Volgograd Region, western Russia. All specimens were recovered in the right slope of the Don River valley from the Lipovskaya Formation. It was named by Sennikov in 2011 and the type species is Augustaburiania vatagini. Augustaburiania is the oldest known tanystropheid.
Youngosuchus is an extinct genus of archosaur from the Middle Triassic of China. The type species is Y. sinensis. Y. sinensis was first described in 1973 as a new species of the erythrosuchid Vjushkovia. In 1985, it was reassigned as its own genus of rauisuchid. A 1992 study supported the original classification of Youngosuchus sinensis as an erythrosuchid, but more recent studies classify it as a "rauisuchian"-grade loricatan archosaur completely unrelated to Vjushkovia, which is most likely a synonym of Garjainia.
Vonhuenia is an extinct genus of proterosuchid, a basal archosauriform from the Early Triassic of Russia. Fossils have been found in the Vokhminskaya Formation, along the Vetluga River that are Induan in age, making Vonhuenia one of the earliest archosauriforms.
Blomosuchus is an extinct genus of archosauriform from the Early Triassic of Russia. The type species was named in 1992 as Blomia georgii. However, the name Blomia was preoccupied by a genus of mites in the family Glycyphagidae (Blomia), so the genus was renamed Blomosuchus in 1997. Fossils of Blomosuchus have been found along the Vetluga River besides fossils of another problematic archosauriform, Vonhuenia.
Exilisuchus is an extinct genus of archosauromorph reptile from the Early Triassic of Russia. The type species E. tubercularis was named in 1979. Fossils are known from two Russian assemblages, the Benthosuchus - Thoosuchus assemblage and the Angusaurus assemblage. Both are Olenekian in age. Although initially believed to be a proterosuchid archosauriform, a 2016 analysis found that it also shared features with tanystropheids, and thus is currently classified as Archosauromorpha incertae sedis. Due to the small amount of preserved material, this genus is sometimes considered a nomen dubium.
Silphedosuchus is an extinct genus of therocephalian therapsids from the Early Triassic of Russia. It is a member of the family Ericiolacertidae, along with the genus Ericiolacerta from South Africa and Antarctica. The type species Silphedosuchus orenburgensis was described in 1977 on the basis of a single holotype skull from Orenburg Oblast.
Coelodontognathus is an extinct genus of reptile from the Early Triassic of European Russia. It was originally described as a procolophonid parareptile in 1967 but was reclassified as a possible trilophosaurid archosauromorph in 2008. The genus includes two species: the type species C. donensis and C. ricovi. C. donensis is known from the holotype PIN 4173/129 and the referred PIN 4173/130, and C. ricovi is known from the holotype PIN 4173/127 and the referred PIN 4173/128, all of which represent dentaries that are housed at the Paleontological Institute, Russian Academy of Sciences. Another dentary, SGU 104/3105, originally referred to C. donensis was reassigned to its own genus and species Vitalia grata by Ivakhnenko, 1973. The fossils have been found at the Donskaya Luka Locality near the village of Sirotinskaya and the Don River in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. Like Coelodontognathus, Vitalia which is known from the same locality was also first identified as a procolophonid and later reclassified as a trilophosaurid. Coelodontognathus and Vitalia are similar to procolophonids in that they have wide teeth but differs from them in that they have tooth roots set deep into the jaws.
Uralosaurus is an extinct genus of erythrosuchid archosauriform known from the Middle Triassic Donguz Formation of southeastern European Russia. It contains a single species, Uralosaurus magnus. It was named by Vitalii Georgievich Ochev in 1980 as a species of Erythrosuchus otherwise known from the Triassic of Africa and reassigned to its own genus by Andrey G. Sennikov in 1995.
Doniceps is an extinct genus of reptile from the Early Triassic of European Russia known from the type species D. lipovensis. It was solely known from the holotype premaxilla 104/3106 housed at Saratov University, however the specimen is probably lost. It was collected at the Donskaya Luka Locality near the village of Sirotinskaya in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. The generic name refers to the nearby Don River added the Greek suffix for "head", -ceps. The specific name is derived from the name of the type locality Donskaya Luka, also known as Lipovaya Balka. Named by Otschev and Rykov in 1968 as an archosauromorph, Arkhangelskii & Sennikov (2008) classified the taxon as a possible trilophosaurid. Doniceps is thought to be similar to Coelodontognathus and Vitalia, both of which are known exclusively from the same locality and were first identified as procolophonids but later reclassified as trilophosaurids.
Vitalia is an extinct genus of reptile from the Early Triassic of European Russia known from the type species V. grata. It is known from the holotype dentary PIN 4173/126 as well as two additional dentaries PIN 1043/627 and 1043/628, all housed at the Paleontological Institute, Russian Academy of Sciences. The type dentary was originally included in the hypodigm of Coelodontognathus donensis named by the notable Russian vertebrate paleontologist Vitaliy Georgiyevich Ochev in 1967. Ivakhnenko (1973) separated the specimen and gave it its own genus and species name in light of the new material, which he named in honor of Ochev. The dentaries of Vitalia were collected at the Donskaya Luka Locality near the village of Sirotinskaya in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. Like Coelodontognathus, Vitalia was originally described as a procolophonid parareptile in 1973, but Arkhangelskii & Sennikov (2008) reclassified the taxon as a possible trilophosaurid archosauromorph. Vitalia is thought to be similar to the possible trilophosaurids Coelodontognathus and Doniceps, both of which are known exclusively from the same locality. Coelodontognathus and Vitalia are similar to procolophonids in that they have wide teeth but differs from them in that they have tooth roots set deep into the jaws.
Megacnemus is an extinct genus of archosauromorph reptile. Megacnemus is an enigmatic and poorly described genus, known from a single bone. The type species of Megacnemus, Megacnemus grandis, was named by Friedrich von Huene in 1954. The holotype of Megacnemus is a limb bone, believed to be a femur. Although the exact locality from which this bone was unearthed is unknown, it is believed to have come from Middle Triassic deposits near Gogolin, in southwest Poland. As the bone is more than 20 centimeters long, Megacnemus may have been quite large, similar to Vritraminosaurus in size.