Guchengosuchus Temporal range: Early Triassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Clade: | Archosauriformes |
Family: | † Erythrosuchidae |
Genus: | † Guchengosuchus Peng, 1991 |
Type species | |
†Guchengosuchus shiguaiensis Peng, 1991 |
Guchengosuchus is an extinct genus of erythrosuchid archosauriform from the Early Triassic of China. It is known from a single holotype skeleton called IVPP V 8808, described in 1991 from the lower Ermaying Formation in Shanxi. The lower Ermaying Formation dates back to the Olenekian stage of the Early Triassic, making Guchengosuchus one of the earliest archosauriforms. [1] IVPP V 8808 is a disarticulated skeleton including a partial skull, lower jaw, some vertebrae, a scapula, and forelimb bones. Like some other erythrosuchids, Guchengosuchus has a tall skull with a notch between the premaxilla and maxilla bones of the upper jaw. The ribs of Guchengosuchus each have three heads, a feature also seen in the Russian erythrosuchid Vjushkovia . [2] When it was first described, Guchengosuchus was placed in the suborder Proterosuchia, a group that included the families Erythrosuchidae, Proterosuchidae, and Proterochampsidae and was thought to be closely related to a primitive group of reptiles called Eosuchia. Proterosuchia is now considered a paraphyletic assemblage of basal archosauriforms representing a series of successive branches of stem group archosaurs.
Erythrosuchidae are a family of large basal archosauriform carnivores that lived from the later Early Triassic (Olenekian) to the early Middle Triassic (Anisian).
Shansisuchus is an extinct genus of archosauriform reptile belonging to the family Erythrosuchidae that lived during the Middle Triassic in what is now China. The first fossils of Shansisuchus were discovered from the Ermaying Formation of Shanxi (Shansi) province in 1964 by Chinese paleontologist Yang Zhongjian. Like other erythrosuchids, Shansisuchus was a large-bodied carnivore with a large, deep skull. Shansisuchus is unique among early archosauriforms in having a hole in its skull called a subnarial fenestra.
Garjainia is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.5–2 metres (4.9–6.6 ft) long. It contained two species, Garjainia prima from the Yarengian/Yarkenskian Supergorizont of Russia, and Garjainia madiba from the Burgersdorp Formation of South Africa. "Vjuskovia triplicostata", a name assigned to some erythrosuchid fossils from Russia, has been synonymized with Garjainia prima.
Proterosuchus is an extinct genus of archosauriform reptiles that lived during the Early Triassic. It contains three valid species: the type species P. fergusi and the referred species P. alexanderi and P. goweri. All three species lived in what is now South Africa. The genus was named in 1903 by the South African paleontologist Robert Broom. The genus Chasmatosaurus is a junior synonym of Proterosuchus.
Doswellia is an extinct genus of archosauriform from the Late Triassic of North America. It is the most notable member of the family Doswelliidae, related to the proterochampsids. Doswellia was a low and heavily built carnivore which lived during the Carnian stage of the Late Triassic. It possesses many unusual features including a wide, flattened head with narrow jaws and a box-like rib cage surrounded by many rows of bony plates. The type species Doswellia kaltenbachi was named in 1980 from fossils found within the Vinita member of the Doswell Formation in Virginia. The formation, which is found in the Taylorsville Basin, is part of the larger Newark Supergroup. Doswellia is named after Doswell, the town from which much of the taxon's remains have been found. A second species, D. sixmilensis, was described in 2012 from the Bluewater Creek Formation of the Chinle Group in New Mexico; however, this species was subsequently transferred to a separate doswelliid genus, Rugarhynchos. Bonafide Doswellia kaltenbachi fossils are also known from the Chinle Formation of Arizona.
Turfanosuchus is a genus of archosauriform reptile, likely a gracilisuchid archosaur, which lived during the Middle Triassic (Anisian) of northwestern China. The type species, T. dabanensis, was described by C.C. Young in 1973, based on a partially complete but disarticulated fossil skeleton found in the Kelamayi Formation of the Turfan Basin.
Fenhosuchus is an extinct genus of archosauriform. The holotype, IVPP V 2697, and referred materials have been found in the Hsishihwa locality at Wuhsiang, China, from the Upper Ermaying Formation. The locality dates back to the Anisian stage of the Middle Triassic. The genus was named after the Fen River in Shanxi Province from which specimens were found. It may prove to be a chimera being composed of material from several different animals. Some material were believed to represent a rauisuchid. The calcaneum of Fenhosuchus seems to belong to an erythrosuchid or other basal archosauriform. Much of the material of the tarsal bones seem to be similar to those of the genus Shansisuchus. According to Nesbitt (2009) the assessment of Gower (2000) was correct, the holotype is a mix of Shansisuchus remains and a possible fragment from a paracrocodylomorph or a dinosauriform. Thus, Fenhosuchus cannot be considered a rauisuchian.
Fugusuchus is an extinct genus of archosauriform, probably the basal-most member of the family Erythrosuchidae. The genus is known from a single fossil from the middle Early Triassic Heshanggou Formation in Shanxi, China. The partial skeleton consists of an incomplete skull, parts of the right forelimb, and an intercentrum. The skeleton, known as GMB V 313, is currently in the Geological Museum of China in Beijing.
Yonghesuchus is an extinct genus of Late Triassic archosaur reptile. Remains have been found from the early Late Triassic Tongchuan Formation in Shanxi, China. It is named after Yonghe County, the county where fossils were found. Currently only one species, Y. sangbiensis, is known. The specific name refers to Sangbi Creek, as fossils were found in one of its banks.
Koilamasuchus is an extinct genus of indeterminate archosauriform from the Triassic of Argentina. It is based on an external mold of a partial postcranial skeleton from the Quebrada de los Fósiles Formation. Due to its incomplete nature, the relationships of this reptile are difficult to establish. Originally described as a non-archosaur archosauriform, later studies tentatively considered it a doswelliid or a suchian archosaur.
Youngosuchus is an extinct genus of archosaur from the Middle Triassic of China. The type species is Y. sinensis. Y. sinensis was first described in 1973 as a new species of the erythrosuchid Vjushkovia. In 1985, it was reassigned as its own genus of rauisuchid. A 1992 study supported the original classification of Youngosuchus sinensis as an erythrosuchid, but more recent studies classify it as a "rauisuchian"-grade loricatan archosaur completely unrelated to Vjushkovia, which is most likely a synonym of Garjainia.
Shaanbeikannemeyeria is an extinct genus of dicynodont known from the Early Triassic of China. It contains a single species, S. xilougoensis, which was described in 1980 by Zheng-Wu Cheng from a skull catalogued as IGCAGS V315. The specimen was lost, and a neotype skull IVPP V 11674 was later designated. A second species, S. buergondia, was named by Jin-Lin Li in 1980 from a partial skeleton, but it has since been regarded as a synonym of S. xilougoensis.
Acallosuchus is an extinct genus of reptile from the Triassic Chinle Formation of the southwestern United States. Although it was discovered in 1923, Acallosuchus was not described until 1995, when the type species A. rectori was named. The taxonomy classification of Acallosuchus is uncertain. Although it is known to be a neodiapsid reptile, it has not been assigned with confidence to any particular group of neodiapsids.
Eumetabolodon is an extinct genus of procolophonine procolophonid parareptile from early and middle Triassic deposits of Nei Mongol, northern China. Two species of Eumetabolodon were named by J. L. Li in 1983 and the type species is Eumetabolodon bathycephalus.
Ordosiodon is an extinct genus of therocephalian therapsids from the Early Triassic of China. It includes two species, O. lincheyuensis and O. youngi.
Wangisuchus is an extinct genus of archosauriform reptile from the Middle Triassic of China that is known from fragmentary fossil jaw bones. These bones were found at the Hsishihwa locality in the upper Ermaying Formation, which dates to the late Anisian stage about 242 million years ago. Wangisuchus was named in 1964 by Chinese paleontologist Yang Zhongjian, who described a single species, Wangisuchus tzeyii, on the basis of these bones.
Asperoris is an extinct genus of archosauriform reptile known from the Middle Triassic Manda Beds of southwestern Tanzania. It is the first archosauriform known from the Manda Beds that is not an archosaur. However, its relationships with other non-archosaurian archosauriforms are uncertain. It was first named by Sterling J. Nesbitt, Richard J. Butler and David J. Gower in 2013 and the type species is Asperoris mnyama. Asperoris means "rough face" in Latin, referring to the distinctive rough texture of its skull bones.
The Ermaying Formation is a geological formation of Anisian age in north-central China. It is found across much of the Ordos Basin, at outcrops within the provinces of Shaanxi, Shanxi, and Inner Mongolia. It is composed of up to 600 m thick sequence of mudstone and sandstone, overlying the Heshanggou Formation and underlying the Tongchuan Formation. In the southern part of the Ordos Basin, the Zhifang Formation is equivalent to the Ermaying Formation.
Kwangsisaurus is an extinct genus of a basal pistosauroid known from the Early or Middle Triassic of Guangxi, southern China. It contains a single species, Kwangsisaurus orientalis.
Polymorphodon is an extinct genus of archosauriform reptile from the Middle Triassic of Germany. The only known species is Polymorphodon adorfi, discovered in Lower Keuper deposits at a quarry in Eschenau, Germany. Polymorphodon is notable for its heterodont dentition, with long and conical premaxillary teeth followed by thin maxillary teeth with large serrations. Maxillary teeth near the back of the mouth are short and leaf-shaped, similar to some living and extinct reptiles with a herbivorous or omnivorous diet. This may suggest that Polymorphodon had some reliance on plants in its diet, a rarity among basal archosauriforms, most of which are carnivores.