Xiaowa Formation

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Xiaowa Formation
Stratigraphic range: Carnian
Type Geological formation
Underlies Laishike Formation
Overlies Zhuganpo Formation
Lithology
Primary limestone, marl
Location
Country China
Extent Yunnan–Guizhou Plateau

The Xiaowa Formation is a Carnian-age geological formation found in southern China. It is a sequence of limestone and marls from the Carnian stage of the Triassic. Its lower section was previously known as the Wayao Formation or Wayao Member of the Falang Formation (a nomenclature still used by some authors). In 2002, the Wayao Member was renamed and raised to the Xiaowa Formation to prevent confusion with an Eocene unit of the same name. [1] Crinoids and marine reptiles are abundant in the Xiaowa Formation, forming a lagerstätte known as the Guanling biota. Ammonoids and conodonts found in the formation constrain its age to the early Carnian. [2] [1] [3] [4] [5] Reptiles of the Guanling biota include ichthyosaurs, thalattosaurs, placodonts, and Odontochelys (an early relative of turtles). [4] Sedimentary events within this formation have been tied to the Carnian Pluvial Event. [6] [7]

Contents

Geology

The Xiaowa Formation has three members. The lower member is relatively thin but is also very fossiliferous. It begins with thick-bedded grey biomicrite (fine-grained fossiliferous limestone) interbedded with greenish shale. Bivalves and crinoid fragments are the most common fossils in the biomicrite layers, which sometimes grade upwards to dark grey laminated marls. The lower section of the lower member represents a relatively well-oxygenated pelagic environment. The lower member then transitions to a section of darker and more clastic layers indicative of anoxic conditions and reduced reef activity. Most of the articulated crinoids and vertebrate fossils of the Guanling biota hail from a dark grey micrite at the base of the lower member's upper section. This is followed by dark grey marls and black shale rich in bivalves, ammonoids, and slightly radioactive clay minerals. The lower member concludes with a sequence of dark grey laminated marls incorporating conodont fossils and silty quartz grains. [1] [4]

The middle member of the Xiaowa Formation is by far the thickest unit and includes thick-bedded grey limestone and marl layers interbedding with each other. Sediment deformation is characteristic of layers in this member, while fossils are represented mainly by occasional bivalves and ammonoids. The middle member represents a deep-water environment influenced by tectonic events which disturb sediment layers and create distant turbidites that periodically supply increased clastic material. The upper member is mostly dominated by laminated limestone. Though marl interbedding and fossils are practically absent, silty to sandy quartz grains are common and dominate the last few meters of the formation. This member represents a shallower ocean environment (likely raised by tectonic uplift) supplied with dust from terrestrial areas [1] [4]

Paleobiota

The Xiaowa Formation encompasses several biostratigraphic zones. The Protrachyceras costulatum ammonoid zone of the upper Zhuganpo Formation continues into the first few meters of the Xiaowa Formation's lower member. However, the rest of the lower member (including the Guanling biota) belongs to the Trachyceras multituberculatum ammonoid zone. This unit has also been called the Austrotrachyceras triadicum zone, and is likely equivalent to the T. aon or T. aonoides zone of the western Tethys (Europe). The middle member of the Xiaowa Formation belongs to the Sirenites cf. senticosus ammonoid zone. [2] [1] [3] [5]

Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Bivalves

Bivalves of the Xiaowa Formation
TaxonSpeciesNotes
Angustella A. sp.A rare bakevelliid bivalve. [1]
Asoella A. sp.A rare asoelliid bivalve. [1]
Daonella
D. bifurcataAn abundant halobiid bivalve, namesake of the H. subcomata-D.bifurcata bivalve zone. [1]
D. bulogensisA halobiid bivalve. [8]
D. indicaA common halobiid bivalve. [1]
Halobia
H. brachyotisA common halobiid bivalve. [1]
H. kuiA common halobiid bivalve. [1]
H. planicostaA common halobiid bivalve. [1]
H. rugosoidesA common halobiid bivalve. [1]
H. subcomataAn abundant halobiid bivalve, namesake of the H. subcomata-D.bifurcata bivalve zone. [1]
Krumbeckiella K. sp.A rare pergamidiid bivalve. [1]
Plagiostoma P. sp.A rare limid bivalve. [1]

Brachiopods

Brachiopods of the Xiaowa Formation
TaxonSpeciesNotes
Crania?sp.A possible craniid brachiopod. [1]
Similingula S. cf. lipoldiA rare brachiopod. [1]

Cephalopods

Cephalopods of the Xiaowa Formation
TaxonSpeciesNotes
Arctosirenites A. canadensisA trachyceratid ammonoid which occurs alongside T. multituberculatum. [2]
A. columbianusA trachyceratid ammonoid which occurs alongside T. multituberculatum. [2]
Austrotrachyceras A. triadicumA trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] [5]
Buchites B cf. aldrovandiiA buchitid ammonoid which occurs in the S. cf. senticosus zone. [1] [4]
Clionites C. cf. zeilleriA clionitid ammonoid which occurs in the P. costulatum zone. [1] [4]
Enoploceras?E. sp.A rare tainoceratid nautiloid tentatively referred to Enoploceras. [2]
Guanlingoceras G. guanlingensisA trachyceratid ammonoid [9]
Hauerites H. cf. himalayanusA rare ammonoid which occurs in the T. multituberculatum zone. [1] [4]
Paratrachyceras P. cf. hoffmaniA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4]
Protrachyceras P. costulatumA trachyceratid ammonoid, namesake of the P. costulatum zone [2] [1] [4]
P. depratiA trachyceratid ammonoid which occurs in the oldest part of the Lower Member. [1] [4]
P. douvilleiA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4]
P. cf. douvilleiA trachyceratid ammonoid which occurs in the P. costulatum zone. [1] [4]
P. ladinumA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1]
P. longiangenseA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1]
P. spAn unnamed species of trachyceratid ammonoid which occurs in the S. cf. senticosus zone. [2] [1] [4]
Simonyceras S. simonyiA ussuritid ammonoid. [3]
Sirenites S. cf. senticosusA trachyceratid ammonoid, namesake of the S. cf. senticosus zone. [1] [4]
Sibyllites S. cf. tenuispinosusA tropitid ammonoid which occurs in the S. cf. senticosus zone. [1]
S. sp.An unnamed species of tropitid ammonoid which occurs in the S. cf. senticosus zone. [1]
Trachyceras T. aonoidesA trachyceratid ammonoid, namesake of the T. aonoides zone. [3]
T. cf. aonA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4]
T. multituberculatumAn abundant trachyceratid ammonoid, namesake of the T. multituberculatum zone. [2] [1] [4] [5]
T. sinensisA trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1]
T. uraniaeA trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] [5]
T. sp. AAn unnamed species of trachyceratid ammonoid similar to Protrachyceras douvillei. [2]
T. sp. BAn unnamed species of trachyceratid ammonoid similar to Protrachyceras deprati. [2]
T. sp.An unnamed species of trachyceratid ammonoid which occurs in the S. cf. senticosus zone. [1] [4]

Conodonts

Conodonts of the Xiaowa Formation
TaxonSpecies
Metapolygnathus / Paragondolella / Quadralellaauriformis [5]
carpathica [10]
foliata foliata [11] [5]
foliata inclinata [11] [5]
jiangyouensis [11]
maantangensis [11] [10]
navicula navicula [11]
nodosus [1] [4]
polygnathiformis [1] [4]
prelindae [12]
robusta [10]
tadpole [11] [5]
wayaoensis [13] [12] [14]
xinpuensis [13]

Echinoderms

Echinoderms of the Xiaowa Formation
TaxonSpeciesNotes
Calclamnidae indet.Holothurian (sea cucumber) ossicles. [1]
Osteocrinus O. cf. spinosusA free-swimming roveacrinid crinoid. [1]
O. cf. virgatusA free-swimming roveacrinid crinoid. [1]
Traumatocrinus T. cf. lipoldiAn abundant traumatocrinid crinoid. Pseudoplanktonic, forming colonies on floating driftwood. [1]

Fish

Fish of the Xiaowa Formation
TaxonSpeciesNotes
Annulicorona A. pyramidalis Elasmobranch denticles [1]
Asialepidotus A. sp.A halecomorph holostean [1]
Birgeria
B. guizhouensisA large predatory birgeriid [4]
B. sp.A large predatory birgeriid [4]
Colobodus C. sp.A colobodontid perleidiform [4]
Guizhoucoelacanthus G. largusA whiteiid coelacanth [4]
Guizhoueugnathus G. largusA eugnathid holostean [4]
Parvicorona P. dacrysulcaElasmobranch denticles [1]
Peltopleurus P. brachycephalusA peltopleurid holostean [1] [4]
Pholidopleurus P. xiaowaensisA pholidopleurid holostean [4]
Saurichthys S. taotieA saurichthyid saurichthyiform [15]

Reptiles

Reptiles of the Xiaowa Formation
TaxonSpeciesNotes
Anshunsaurus A. huangguoshuensisA large askeptosauroid thalattosaur [4]
"Callawayia""C." wolonggangensisA shastasaurid ichthyosaur. Originally referred to Callawayia , but likely unrelated. [16]
Concavispina C. biseridensA large thalattosauroid thalattosaur [17]
Cyamodus C. orientalisA cyamodontid placodont [18]

"Cymbospondylus"

"C." asiaticus

A junior synonym of Guizhouichthyosaurus tangae, unrelated to Cymbospondylus [4]

Eorhynchochelys E. sinensisA large aquatic pantestudine (early turtle relative) [19]
Guanlingsaurus G. liangaeA giant shastasaurid ichthyosaur [4]
Guizhouichthyosaurus G. tangaeA common shastasaurid ichthyosaur [4]
Miodentosaurus M. brevisA large askeptosauroid thalattosaur [4]
Neosinasaurus N. hoangiA poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20]
Odontochelys O. semitestaceaAn aquatic pantestudine [21] [19]

Panjiangsaurus

P. epicharis

A junior synonym of Guizhouichthyosaurus tangae [4]

Psephochelys P. polyosteodermaA placochelyid placodont [4]
Qianichthyosaurus Q. zhouiA common small toretocnemid ichthyosaur [4]
Sinocyamodus S. xinpuensisA cyamodontoid placodont [4]

Typicusichthyosaurus

T. tsaihuae

Likely a junior synonym of Guanlingsaurus lingae [4]

Wayaosaurus W. bellusA poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20]
W. geeiA poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20]
Xinpusaurus

X. bamaolinensis

A thalattosauroid thalattosaur, [4] possibly an invalid synonym of another Xinpusaurus species [22] [23]

X. kohiA thalattosauroid thalattosaur [23]
X. suniA thalattosauroid thalattosaur [4]

References

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