Xiaowa Formation | |
---|---|
Stratigraphic range: Carnian | |
Type | Geological formation |
Underlies | Laishike Formation |
Overlies | Zhuganpo Formation |
Lithology | |
Primary | limestone, marl |
Location | |
Country | China |
Extent | Yunnan–Guizhou Plateau |
The Xiaowa Formation is a Carnian-age geological formation found in southern China. It is a sequence of limestone and marls from the Carnian stage of the Triassic. Its lower section was previously known as the Wayao Formation or Wayao Member of the Falang Formation (a nomenclature still used by some authors). In 2002, the Wayao Member was renamed and raised to the Xiaowa Formation to prevent confusion with an Eocene unit of the same name. [1] Crinoids and marine reptiles are abundant in the Xiaowa Formation, forming a lagerstätte known as the Guanling biota. Ammonoids and conodonts found in the formation constrain its age to the early Carnian. [2] [1] [3] [4] [5] Reptiles of the Guanling biota include ichthyosaurs, thalattosaurs, placodonts, and Odontochelys (an early relative of turtles). [4] Sedimentary events within this formation have been tied to the Carnian Pluvial Event. [6] [7]
The Xiaowa Formation has three members. The lower member is relatively thin but is also very fossiliferous. It begins with thick-bedded grey biomicrite (fine-grained fossiliferous limestone) interbedded with greenish shale. Bivalves and crinoid fragments are the most common fossils in the biomicrite layers, which sometimes grade upwards to dark grey laminated marls. The lower section of the lower member represents a relatively well-oxygenated pelagic environment. The lower member then transitions to a section of darker and more clastic layers indicative of anoxic conditions and reduced reef activity. Most of the articulated crinoids and vertebrate fossils of the Guanling biota hail from a dark grey micrite at the base of the lower member's upper section. This is followed by dark grey marls and black shale rich in bivalves, ammonoids, and slightly radioactive clay minerals. The lower member concludes with a sequence of dark grey laminated marls incorporating conodont fossils and silty quartz grains. [1] [4]
The middle member of the Xiaowa Formation is by far the thickest unit and includes thick-bedded grey limestone and marl layers interbedding with each other. Sediment deformation is characteristic of layers in this member, while fossils are represented mainly by occasional bivalves and ammonoids. The middle member represents a deep-water environment influenced by tectonic events which disturb sediment layers and create distant turbidites that periodically supply increased clastic material. The upper member is mostly dominated by laminated limestone. Though marl interbedding and fossils are practically absent, silty to sandy quartz grains are common and dominate the last few meters of the formation. This member represents a shallower ocean environment (likely raised by tectonic uplift) supplied with dust from terrestrial areas [1] [4]
The Xiaowa Formation encompasses several biostratigraphic zones. The Protrachyceras costulatum ammonoid zone of the upper Zhuganpo Formation continues into the first few meters of the Xiaowa Formation's lower member. However, the rest of the lower member (including the Guanling biota) belongs to the Trachyceras multituberculatum ammonoid zone. This unit has also been called the Austrotrachyceras triadicum zone, and is likely equivalent to the T. aon or T. aonoides zone of the western Tethys (Europe). The middle member of the Xiaowa Formation belongs to the Sirenites cf. senticosus ammonoid zone. [2] [1] [3] [5]
Color key
| Notes Uncertain or tentative taxa are in small text; |
Bivalves of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Angustella | A. sp. | A rare bakevelliid bivalve. [1] |
Asoella | A. sp. | A rare asoelliid bivalve. [1] |
Daonella | ||
D. bifurcata | An abundant halobiid bivalve, namesake of the H. subcomata-D.bifurcata bivalve zone. [1] | |
D. bulogensis | A halobiid bivalve. [8] | |
D. indica | A common halobiid bivalve. [1] | |
Halobia | ||
H. brachyotis | A common halobiid bivalve. [1] | |
H. kui | A common halobiid bivalve. [1] | |
H. planicosta | A common halobiid bivalve. [1] | |
H. rugosoides | A common halobiid bivalve. [1] | |
H. subcomata | An abundant halobiid bivalve, namesake of the H. subcomata-D.bifurcata bivalve zone. [1] | |
Krumbeckiella | K. sp. | A rare pergamidiid bivalve. [1] |
Plagiostoma | P. sp. | A rare limid bivalve. [1] |
Brachiopods of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Crania? | sp. | A possible craniid brachiopod. [1] |
Similingula | S. cf. lipoldi | A rare brachiopod. [1] |
Cephalopods of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Arctosirenites | A. canadensis | A trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] |
A. columbianus | A trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] | |
Austrotrachyceras | A. triadicum | A trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] [5] |
Buchites | B cf. aldrovandii | A buchitid ammonoid which occurs in the S. cf. senticosus zone. [1] [4] |
Clionites | C. cf. zeilleri | A clionitid ammonoid which occurs in the P. costulatum zone. [1] [4] |
Enoploceras? | E. sp. | A rare tainoceratid nautiloid tentatively referred to Enoploceras. [2] |
Guanlingoceras | G. guanlingensis | A trachyceratid ammonoid [9] |
Hauerites | H. cf. himalayanus | A rare ammonoid which occurs in the T. multituberculatum zone. [1] [4] |
Paratrachyceras | P. cf. hoffmani | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4] |
Protrachyceras | P. costulatum | A trachyceratid ammonoid, namesake of the P. costulatum zone [2] [1] [4] |
P. deprati | A trachyceratid ammonoid which occurs in the oldest part of the Lower Member. [1] [4] | |
P. douvillei | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4] | |
P. cf. douvillei | A trachyceratid ammonoid which occurs in the P. costulatum zone. [1] [4] | |
P. ladinum | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] | |
P. longiangense | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] | |
P. sp | An unnamed species of trachyceratid ammonoid which occurs in the S. cf. senticosus zone. [2] [1] [4] | |
Simonyceras | S. simonyi | A ussuritid ammonoid. [3] |
Sirenites | S. cf. senticosus | A trachyceratid ammonoid, namesake of the S. cf. senticosus zone. [1] [4] |
Sibyllites | S. cf. tenuispinosus | A tropitid ammonoid which occurs in the S. cf. senticosus zone. [1] |
S. sp. | An unnamed species of tropitid ammonoid which occurs in the S. cf. senticosus zone. [1] | |
Trachyceras | T. aonoides | A trachyceratid ammonoid, namesake of the T. aonoides zone. [3] |
T. cf. aon | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] [4] | |
T. multituberculatum | An abundant trachyceratid ammonoid, namesake of the T. multituberculatum zone. [2] [1] [4] [5] | |
T. sinensis | A trachyceratid ammonoid which occurs in the T. multituberculatum zone. [1] | |
T. uraniae | A trachyceratid ammonoid which occurs alongside T. multituberculatum. [2] [5] | |
T. sp. A | An unnamed species of trachyceratid ammonoid similar to Protrachyceras douvillei. [2] | |
T. sp. B | An unnamed species of trachyceratid ammonoid similar to Protrachyceras deprati. [2] | |
T. sp. | An unnamed species of trachyceratid ammonoid which occurs in the S. cf. senticosus zone. [1] [4] | |
Conodonts of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | |
Metapolygnathus / Paragondolella / Quadralella | auriformis [5] | |
carpathica [10] | ||
foliata foliata [11] [5] | ||
foliata inclinata [11] [5] | ||
jiangyouensis [11] | ||
maantangensis [11] [10] | ||
navicula navicula [11] | ||
nodosus [1] [4] | ||
polygnathiformis [1] [4] | ||
prelindae [12] | ||
robusta [10] | ||
tadpole [11] [5] | ||
wayaoensis [13] [12] [14] | ||
xinpuensis [13] | ||
Echinoderms of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Calclamnidae | indet. | Holothurian (sea cucumber) ossicles. [1] |
Osteocrinus | O. cf. spinosus | A free-swimming roveacrinid crinoid. [1] |
O. cf. virgatus | A free-swimming roveacrinid crinoid. [1] | |
Traumatocrinus | T. cf. lipoldi | An abundant traumatocrinid crinoid. Pseudoplanktonic, forming colonies on floating driftwood. [1] |
Fish of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Annulicorona | A. pyramidalis | Elasmobranch denticles [1] |
Asialepidotus | A. sp. | A halecomorph holostean [1] |
Birgeria | ||
B. guizhouensis | A large predatory birgeriid [4] | |
B. sp. | A large predatory birgeriid [4] | |
Colobodus | C. sp. | A colobodontid perleidiform [4] |
Guizhoucoelacanthus | G. largus | A whiteiid coelacanth [4] |
Guizhoueugnathus | G. largus | A eugnathid holostean [4] |
Parvicorona | P. dacrysulca | Elasmobranch denticles [1] |
Peltopleurus | P. brachycephalus | A peltopleurid holostean [1] [4] |
Pholidopleurus | P. xiaowaensis | A pholidopleurid holostean [4] |
Saurichthys | S. taotie | A saurichthyid saurichthyiform [15] |
Reptiles of the Xiaowa Formation | ||
---|---|---|
Taxon | Species | Notes |
Anshunsaurus | A. huangguoshuensis | A large askeptosauroid thalattosaur [4] |
"Callawayia" | "C." wolonggangensis | A shastasaurid ichthyosaur. Originally referred to Callawayia , but likely unrelated. [16] |
Concavispina | C. biseridens | A large thalattosauroid thalattosaur [17] |
Cyamodus | C. orientalis | A cyamodontid placodont [18] |
"Cymbospondylus" | "C." asiaticus | A junior synonym of Guizhouichthyosaurus tangae, unrelated to Cymbospondylus [4] |
Eorhynchochelys | E. sinensis | A large aquatic pantestudine (early turtle relative) [19] |
Guanlingsaurus | G. liangae | A giant shastasaurid ichthyosaur [4] |
Guizhouichthyosaurus | G. tangae | A common shastasaurid ichthyosaur [4] |
Miodentosaurus | M. brevis | A large askeptosauroid thalattosaur [4] |
Neosinasaurus | N. hoangi | A poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20] |
Odontochelys | O. semitestacea | An aquatic pantestudine [21] [19] |
Panjiangsaurus | P. epicharis | A junior synonym of Guizhouichthyosaurus tangae [4] |
Psephochelys | P. polyosteoderma | A placochelyid placodont [4] |
Qianichthyosaurus | Q. zhoui | A common small toretocnemid ichthyosaur [4] |
Sinocyamodus | S. xinpuensis | A cyamodontoid placodont [4] |
Typicusichthyosaurus | T. tsaihuae | Likely a junior synonym of Guanlingsaurus lingae [4] |
Wayaosaurus | W. bellus | A poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20] |
W. geei | A poorly-known marine reptile. Originally described as a pachypleurosaur, though may be a thalattosaur instead. [20] | |
Xinpusaurus | X. bamaolinensis | A thalattosauroid thalattosaur, [4] possibly an invalid synonym of another Xinpusaurus species [22] [23] |
X. kohi | A thalattosauroid thalattosaur [23] | |
X. suni | A thalattosauroid thalattosaur [4] | |
In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between 251.2 Ma and 247.2 Ma. The Olenekian is sometimes divided into the Smithian and the Spathian subages or substages. The Olenekian follows the Induan and is followed by the Anisian.
Thalattosauria is an extinct order of marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.
Miodentosaurus is a genus of thalattosaurian from the Late Triassic of China. It is one of several thalattosaurs found in the Xiaowa Formation, also known as the Wayao Member of the Falang Formation. The genus name "Miodentosaurus" translates to "Few toothed-lizard" while the species name "brevis" means "short", in reference to its short snout.
Colobodus is an extinct genus of marine Triassic ray-finned fish of the family Colobodontidae and order Perleidiformes. Fossils have been found in Europe and China, encompassing the former Tethys Ocean. It could reach body lengths of about 70 cm.
Guizhouichthyosaurus is an extinct genus of ichthyosaur which is known primarily from the Xiaowa Formation of the lower Carnian stage of the Late Triassic in southwest China. The type species of this genus is Guizhouichthyosaurus tangae, of which multiple skeletons are known. It has been reassigned as a species of the genus Shastasaurus in the past, though it has since been considered distinct. The ichthyosaurs Cymbospondylus asiaticus, named in 2002, and Panjiangsaurus epicharis, named in 2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia" wollongangense may belong to Guizhouichthyosaurus.
Qianichthyosaurus is an extinct genus of ichthyosaur from the Ladinian and Carnian stages of the Late Triassic epoch. Its fossils have been found in southeastern China, in Carnian rocks of the Falang Formation near Huangtutang, Guizhou. The type species is Qianichthyosaurus zhoui, named by Chun Li in 1999. A second species, Qianichthyosaurus xingyiensis, was named from older (Ladinian) deposits in the Falang Formation in 2013 by Pengfei Yang and colleagues. Complete Qianichthyosaurus fossils are common in the Xiaowa Formation, with both juveniles and pregnant specimens being known; its larger contemporaries, Guizhouichthyosaurus and Guanlingsaurus, are rarer.
The Guanling Formation is a Middle Triassic geologic formation in southwestern China.
Sinosaurichthys is an extinct genus of saurichthyid ray-finned fish, which existed in south-western China during the Middle Triassic. Fossils have been found in the Upper Member of the Guanling Formation of two localities: Yangjuan of Panxian County, Guizhou Province, and Dawazi of Luoping, Yunnan Province, China.
Sinosaurosphargis is an extinct genus of basal marine saurosphargid reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a single species, Sinosaurosphargis yunguiensis.
Anshunsaurus is a genus of thalattosaurs within the family Askeptosauridae. Fossils have been found from Middle Triassic deposits in Guizhou, China. Three species are known: the type species A. huangguoshuensis, the slightly older species A. wushaensis, and the species A. huangnihensis.
Wumengosaurus is an extinct aquatic reptile from the Middle Triassic Guanling Formation of Guizhou, southwestern China. It was originally described as a basal eosauropterygian and usually is recovered as such by phylogenetic analyses, although one phylogeny has placed it as the sister taxon to Ichthyosauromorpha while refraining from a formal re-positioning. It was a relatively small reptile, measuring 95.5–130.5 cm (3.13–4.28 ft) in total body length.
Guanlingsaurus is an extinct genus of shastasaurid ichthyosaur from the Late Triassic of China. It grew up to 8.3 metres (27 ft) in length and has a wide, triangular skull with a short and toothless snout.
Yunguisaurus is an extinct genus of pistosaur known from the Guizhou Province of China.
Sinocyamodus is an extinct genus of placodont reptile from the Late Triassic (Tuvalian) Xiaowa Formation of China. Only one species, Sinocyamodus xinpuensis, is currently assigned to this genus. This genus was the first placodont to be discovered in the eastern Tethys, overthrowing traditional views that the group was restricted to the western Tethys.
Concavispina is an extinct genus of thalattosaur reptile from the early Late Triassic Xiaowa Formation of Guangling, Guizhou, southern China. It contains a single species, Concavispina biseridens. It is known only from the holotype ZMNH M8804, a nearly complete 364 cm long skeleton. Concavispina can be differentiated from other thalattosaurs by possessing two rows of blunt teeth on the anterior part of the maxilla and a V-shaped notch on the dorsal margin of each neural spine in the dorsal (back) vertebrae. Both its generic and specific names refer to these autapomorphies, as Concavispina means "concave spine" and biseridens means "two rows of teeth". It is thought to be most closely related to Xinpusaurus, as both taxa share three derived characters: a maxilla that is curved upward at its anterior end, a humerus that is wider near the shoulder than near the elbow, and the presence of less than five cervicals.
The Carnian pluvial episode (CPE), often called the Carnian pluvial event, was a period of major change in global climate that coincided with significant changes in Earth's biota both in the sea and on land. It occurred during the latter part of the Carnian Stage, a subdivision of the late Triassic period, and lasted for perhaps 1–2 million years.
Xinpusaurus is an extinct genus of thalattosaur from the Late Triassic of Guanling in Guizhou, China. Several species have been named since 2000: the type species X. suni along with the species X. bamaolinensis and X. kohi. A 2013 study proposed that all three species are synonymous with each other, in which case X. suni would be the only valid species, although a 2014 study argued that X. kohi was also valid. A fourth species, X. xingyiensis, was described in 2016.
Saurosphargidae is an extinct family of marine reptiles known from the Early Triassic and early Middle Triassic of Europe and China.
The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.
The Yangliujing Formation is a Middle Triassic geologic unit found in the Guizhou and Yunnan Provinces of southern China.
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