Guanling Formation

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Guanling Formation
Stratigraphic range: early-mid Anisian (Pelsonian)
~247–245  Ma
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Type Geological formation
Underlies Yangliujing Formation
Overlies Jialingjiang Formation
Lithology
Primary Mudstone, limestone
Other Dolomite
Location
Coordinates 25°30′N104°54′E / 25.5°N 104.9°E / 25.5; 104.9
Approximate paleocoordinates 11°42′N94°12′E / 11.7°N 94.2°E / 11.7; 94.2
Region Guizhou & Yunnan Provinces
CountryFlag of the People's Republic of China.svg  China
Extent Yunnan–Guizhou Plateau
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Guanling Formation (China)
China Guizhou location map.jpg
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Guanling Formation (Guizhou)

The Guanling Formation is a Middle Triassic (Anisian or Pelsonian in the regional chronostratigraphy) geologic formation in southwestern China.

Contents

Geology

The formation encompasses two members. The first member is primarily calcareous mudstone and dolomite, indicative of a coastal environment. The second member is a thicker marine sequence of dark micritic limestone with some dolomite. Two distinct fossil assemblages are found in the second member. The older Luoping biota preserves abundant arthropods along with fossils from other invertebrates and vertebrates, which are rare but well-preserved. The slightly younger Panxian fauna has a more diverse and common assortment of marine reptiles such as sauropterygians. [1]

A tuff bed in the Luoping biota has been dated to 244.5 ± 2.2 Ma via U-Pb SHRIMP dating. The Luoping biota as a whole was deposited over a period of 340 ± 71 kyr. This estimate is justified by cyclostratigraphic evidence, as indicated by fluctuations of terrestrial proxy metals. [2]

Fossil content

Among others, the following fossils were reported from the formation:

Ichthyopterygia

Ichthyosauria
GenusSpeciesMemberNotesImage
Barracudasauroides [3] B. panxianensisUpper Member [3] [1] A small mixosaurid ichthyosaur, either a member of its own genus [4] or a species of Mixosaurus . [1]
Phalarodon [5] [6] P. cf. fraasi [5] Upper Member [1] Known from a skull of a small mixosaurid ichthyosaur with durophagous dentition.
P. atavus [6] Upper Member [1] A small mixosaurid ichthyosaur known from a complete skeleton. Might represent a member of its own genus, Contectopalatus .
Xinminosaurus [7] X. catactesUpper Member [1] A medium-sized early ichthyosaur with durophagous dentition, possibly a member of Cymbospondylidae.

Sauropterygia

Placodontiformes
GenusSpeciesMemberNotesImage
Atopodentatus [8] A. unicusUpper Member [8] A bizarre, likely herbivorous [9] sauropterygian with possible placodontiform affinities, [10] known from several well-preserved specimens.
Placodus [11] P. inexpectatusUpper Member [11] [1] A medium-sized placodontid placodont, known from a complete specimen. [11]
Saurosphargidae
GenusSpeciesMemberNotesImage
Largocephalosaurus L. polycarpon [12] Upper Member [12] [1] A mid-sized basal saurosphargid sauropterygian.
L. qianensis [13] Upper Member [13] [1] A mid-sized basal saurosphargid sauropterygian.
Sinosaurosphargis [14] S. yunguinensisUpper Member [14] [1] A small armoured saurosphargid sauropterygian.

Related Research Articles

<i>Dinocephalosaurus</i> Extinct genus of reptiles

Dinocephalosaurus is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, D. orientalis, which was named by Chun Li in 2003. Unlike other long-necked protorosaurs, Dinocephalosaurus convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative Pectodens, which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, Dinocephalosaurus probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as suggested by the poorly-ossified and paddle-like limbs which would have prevented it from going ashore.

<i>Mixosaurus</i> Extinct genus of reptiles

Mixosaurus is an extinct genus of Middle Triassic ichthyosaur. Its fossils have been found near the Italy–Switzerland border and in South China.

<span class="mw-page-title-main">Thalattosauria</span> Extinct group of marine reptiles

Thalattosauria is an extinct order of marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.

<i>Macrocnemus</i> Extinct genus of reptiles

Macrocnemus is an extinct genus of archosauromorph reptile known from the Middle Triassic of Europe and China. Macrocnemus is a member of the Tanystropheidae family and includes three species. Macrocnemus bassanii, the first species to be named and described, is known from the Besano Formation and adjacent paleontological sites in the Italian and Swiss Alps. Macrocnemus fuyuanensis, on the other hand, is known from the Zhuganpo Formation in southern China. A third species, Macrocnemus obristi, is known from the Prosanto Formation of Switzerland and is characterized by gracile limbs. The name Macrocnemus is Greek for "long tibia".

<i>Miodentosaurus</i> Extinct genus of reptiles

Miodentosaurus is a genus of thalattosaurian from the Late Triassic of China. It is one of several thalattosaurs found in the Xiaowa Formation, also known as the Wayao Member of the Falang Formation. The genus name "Miodentosaurus" translates to "Few toothed-lizard" while the species name "brevis" means "short", in reference to its short snout.

<i>Helveticosaurus</i> Extinct genus of reptiles

Helveticosaurus is an extinct genus of diapsid marine reptile known from the Middle Triassic of southern Switzerland. It contains a single species, Helveticosaurus zollingeri, known from the nearly complete holotype T 4352 collected at Cava Tre Fontane of Monte San Giorgio, an area well known for its rich record of marine life during the Middle Triassic.

<i>Sinosaurosphargis</i> Extinct genus of reptiles

Sinosaurosphargis is an extinct genus of basal marine saurosphargid reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a single species, Sinosaurosphargis yunguiensis.

<i>Wumengosaurus</i> Extinct genus of reptiles

Wumengosaurus is an extinct aquatic reptile from the Middle Triassic Guanling Formation of Guizhou, southwestern China. It was originally described as a basal eosauropterygian and usually is recovered as such by phylogenetic analyses, although one phylogeny has placed it as the sister taxon to Ichthyosauromorpha while refraining from a formal re-positioning. It was a relatively small reptile, measuring 95.5–130.5 cm (3.13–4.28 ft) in total body length.

Largocephalosaurus is an extinct genus of basal saurosphargid, a marine reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a type species, Largocephalosaurus polycarpon, and a second species L. qianensis.

<i>Thalattoarchon</i> Extinct genus of reptiles

Thalattoarchon is a genus of large, Middle Triassic predatory ichthyosaur from North America, containing the single species T. saurophagis. The taxon was described in 2013 from a single specimen discovered in the Favret Formation in Nevada. The generic name, meaning "ruler of the seas", refers to its status as an apex predator, while the specific epithet, meaning "lizard eater", alludes to its carnivorous diet. The classification of this genus within the ichthyosaurs is much debated, being either classified within the clade Merriamosauria or in the more basal family Cymbospondylidae.

Eohupehsuchus is a genus of extinct aquatic diapsid from the Upper Spathian of Hubei Province, located in Central China. The genus is monotypic and belongs to the order Hupehsuchia, whose members are characterized by toothless beak-like snouts, a row of dermal plates along their backs, and aquatic adaptations including paddle-shaped limbs and fusiform bodies with pachyostotic ribs.

Eretmorhipis is an extinct genus of hupehsuchian marine reptiles from the Early Triassic of China. It is currently known from two specimens that were discovered in an exposure of the Jialingjiang Formation in Yuan'an County, Hubei, and referred to the newly named species Eretmorhipis carrolldongi in 2015.

<span class="mw-page-title-main">Saurosphargidae</span> Extinct family of reptiles

Saurosphargidae is an extinct family of marine reptiles known from the Early Triassic and early Middle Triassic of Europe and China.

Saurosphargis is an extinct genus of a basal marine reptile, saurosphargid, known from the Middle Triassic of southwestern Poland and eastern Netherlands. It contains a single species, Saurosphargis volzi.

<i>Eusaurosphargis</i> Extinct genus of reptiles

Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi. It was a small reptile, measuring 20 cm (7.9 in) long.

The Xiaowa Formation is a Carnian-age geological formation found in southern China. It is a sequence of limestone and marls from the Carnian stage of the Triassic. Its lower section was previously known as the Wayao Formation or Wayao Member of the Falang Formation. In 2002, the Wayao Member was renamed and raised to the Xiaowa Formation to prevent confusion with an Eocene unit of the same name. Crinoids and marine reptiles are abundant in the Xiaowa Formation, forming a lagerstätte known as the Guanling biota. Ammonoids and conodonts found in the formation constrain its age to the early Carnian. Reptiles of the Guanling biota include ichthyosaurs, thalattosaurs, placodonts, and Odontochelys. Sedimentary events within this formation have been tied to the Carnian Pluvial Event.

The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.

This list of fossil reptiles described in 2014 is a list of new taxa of fossil reptiles that were described during the year 2014, as well as other significant discoveries and events related to reptile paleontology that occurred in 2014.

<i>Luopingosaurus</i> Extinct genus of sauropterygian reptiles

Luopingosaurus is an extinct genus of pachypleurosaurid sauropterygian from the Middle Triassic Guanling Formation of Yunnan Province, China. The genus contains a single species, L. imparilis, known from a well-preserved, nearly complete skeleton.

References

  1. 1 2 3 4 5 6 7 8 9 10 Benton, Michael J.; Zhang, Qiyue; Hu, Shixue; Chen, Zhong-Qiang; Wen, Wen; Liu, Jun; Huang, Jinyuan; Zhou, Changyong; Xie, Tao; Tong, Jinnan; Choo, Brian (2013-10-01). "Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction". Earth-Science Reviews. 125: 199–243. Bibcode:2013ESRv..125..199B. doi:10.1016/j.earscirev.2013.05.014. ISSN   0012-8252.
  2. Ma, Zhixin; Wu, Huaichun; Hu, Shixue; Fang, Qiang; Liu, Xiting; Zhou, Changyong; Wen, Wen; Zhang, Qiyue; Huang, Jinyuan; Min, Xiao (2023-09-26). "Temporal duration and preservation mechanism of the Middle Triassic Luoping biota from South China constrained by geochronology and cyclostratigraphy". Global and Planetary Change. 229: 104254. Bibcode:2023GPC...22904254M. doi:10.1016/j.gloplacha.2023.104254. ISSN   0921-8181. S2CID   263214751.
  3. 1 2 Jiang, Da-Yong; Schmitz, Lars; Hao, Wei-Cheng; Sun, Yuan-Lin (March 30, 2006). "A new mixosaurid ichthyosaur from the Middle Triassic of China". Journal of Vertebrate Paleontology. 26 (1): 60–69. doi:10.1671/0272-4634(2006)26[60:ANMIFT]2.0.CO;2. JSTOR   4524536. S2CID   55171917.
  4. Maisch, Michael W. (2010). "Phylogeny, systematics, and origin of the Ichthyosauria - the state of the art". Palaeodiversity. 3: 141–214.
  5. 1 2 Jiang, Da-Yong; Lars, Schmitz; Motani, Ryosuke; Hao, Wie-Cheng; Sun, Yuan-Lin (2007). "The Mixosaurid Ichthyosaur Phalarodon cf. P. fraasi from the Middle Triassic of Guizhou Province, China". Journal of Paleontology. 81 (3): 602–605. Bibcode:2007JPal...81..602J. doi:10.1666/05092.1. JSTOR   4498810. S2CID   54761545.
  6. 1 2 Liu, Jun; Motani, Ryosuke; Jiang, Da-Yong; Hu, Shi-Xue; Aitchison, Jonathan C.; Rieppel, Olivier; Benton, Michael J.; Zhang, Qi-Yue; Zhou, Chang-Yong (2013). "The first specimen of the Middle Triassic Phalarodon atavus (Ichthyosauria: Mixosauridae) from South China, showing postcranial anatomy and peri-Tethyan distribution". Palaeontology. 56 (4): 849–866. Bibcode:2013Palgy..56..849L. doi:10.1111/pala.12021.
  7. Jiang, Da-Yong; Motani, Ryosuke; Hao, Wei-Cheng; Schmitz, Lars; Rieppel, Olivier; Sun, Yuan-Lin; Sun, Zuo-Yu (2008). "New primitive ichthyosaurian (Reptilia, Diapsida) from the Middle Triassic of Panxian, Guizhou, southwestern China and its position in the Triassic biotic recovery". Progress in Natural Science. 18 (10): 1315–1319. Bibcode:2008PNSMI..18.1315J. doi: 10.1016/j.pnsc.2008.01.039 .
  8. 1 2 Cheng, Long; Chen, Xiao-Hong; Shang, Qing-Hua; Wu, Xiao-Chun (2014). "A new marine reptile from the Triassic of China, with a highly specialized feeding adaptation". Naturwissenschaften. 101 (3): 251–259. Bibcode:2014NW....101..251C. doi:10.1007/s00114-014-1148-4. PMID   24452285. S2CID   7770583.
  9. Chun, Li; Rieppel, Olivier; Long, Cheng; Fraser, Nicholas C. (2016). "The earliest herbivorous marine reptile and its remarkable jaw apparatus". Science Advances. 2 (5): e1501659. Bibcode:2016SciA....2E1659C. doi:10.1126/sciadv.1501659. PMC   4928886 . PMID   27386529.
  10. Wolniewicz, Andrzej S.; Shen, Yuefeng; Li, Qiang; Sun, Yuanyuan; Qiao, Yu; Chen, Yajie; Hu, Yi-Wei; Liu, Jun (2023). "An armoured marine reptile from the Early Triassic of South China and its phylogenetic and evolutionary implications". eLife. 12: e83163. doi: 10.7554/eLife.83163 . PMC   10499374 . PMID   37551884.
  11. 1 2 3 Jiang, Da-Yong; Motani, Ryosuke; Hao, Wei-Cheng; Rieppel, Olivier; Sun, Yuan-Lin; Schmitz, Lars; Sun, Zu-Yu (2008). "First Record of Placodontoidea (Reptilia, Sauropterygia, Placodontia) from the Eastern Tethys". Journal of Vertebrate Paleontology. 28 (3): 904–908. doi:10.1671/0272-4634(2008)28[904:FROPRS]2.0.CO;2. JSTOR   20491013. S2CID   9784064.
  12. 1 2 Cheng, Long; Chen, Xiahong; Zeng, Xiongwei; Cai, Yongjian (2012). "A new eosauropterygian (Diapsida: Sauropterygia) from the Middle Triassic of Luoping, Yunnan Province". Journal of Earth Science. 23 (1): 33–40. Bibcode:2012JEaSc..23...33C. doi:10.1007/s12583-012-0231-z.
  13. 1 2 Li, Chun; Jiang, Da-Yong; Cheng, Long; Wu, Xiao-Chun; Rieppel, Olivier (2014). "A new species of Largocephalosaurus (Diapsida: Saurosphargidae), with implications for the morphological diversity and phylogeny of the group". Geological Magazine. 151 (1): 100–120. Bibcode:2014GeoM..151..100L. doi:10.1017/S001675681300023X.
  14. 1 2 Li, Chun; Rieppel, Olivier; Wu, Xiao-Chun; Zhao, Li-Jun; Wang, Li-Ting (2011). "A new Triassic marine reptile from southwestern China". Journal of Vertebrate Paleontology. 31 (2): 303–312. Bibcode:2011JVPal..31..303L. doi:10.1080/02724634.2011.550368.
  15. Wang, W.; Lei, H.; Li, C. (2023). "A small-sized dinocephalosaurid archosauromorph from the Middle Triassic of Yunnan, southwestern China". Vertebrata PalAsiatica. 62 (1): 13. doi:10.19615/j.cnki.2096-9899.231013.
  16. Xu, G.-H.; Ma, X.-Y. (2023). "A new basal ginglymodian fish (Holostei: Neopterygii) from the Middle Triassic (Anisian) Luoping Biota, Yunnan, China". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlad144.
  17. 1 2 3 Lu, Yu-Ting; Liu, Jun (2023-10-02). "A new tanystropheid (Diapsida: Archosauromorpha) from the Middle Triassic of SW China and the biogeographical origin of Tanystropheidae". Journal of Systematic Palaeontology. 21 (1). Bibcode:2023JSPal..2150778L. doi:10.1080/14772019.2023.2250778. ISSN   1477-2019. S2CID   263634664.
  18. Xu, G.-H.; Shang, Q.-H.; Wang, W.; Ren, Y.; Lei, H.; Liao, J.-L.; Zhao, L.-J.; Li, C. (2023). "A new long-snouted marine reptile from the Middle Triassic of China illuminates pachypleurosauroid evolution". Scientific Reports. 13 (1). 16. Bibcode:2023NatSR..13...16X. doi: 10.1038/s41598-022-24930-y . ISSN   2045-2322. PMC   9816097 . PMID   36604433.