Sinosaurosphargis Temporal range: Middle Triassic, | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Family: | † Saurosphargidae |
Genus: | † Sinosaurosphargis Li et al., 2011 |
Type species | |
†Sinosaurosphargis yunguiensis Li et al., 2011 |
Sinosaurosphargis is an extinct genus of basal marine saurosphargid reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a single species, Sinosaurosphargis yunguiensis. [1]
Saurosphargis is known from several individuals, all of which were collected from Member II of the Guanling Formation, dating to the Pelsonian substage of the latest Anisian stage of the early Middle Triassic, about 243 million years ago. The holotype IVPP V 17040 and paratype IVPP V 16076 are housed at the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, and represent a nearly complete articulated skeleton and skull, and a partial disarticulated postcranial skeleton including dorsal vertebra, ribs, with osteoderms and gastral rib fragments, respectively. ZMNH M 8797, an incomplete postcranial skeleton showing a very well preserved right forelimb housed at the Zhejiang Museum of Nature History of Hangzhou, was also referred to Saurosphargis in its original description. The holotype and ZMNH M 8797 were collected at Yangmazhai of Luoping County, Yunnan Province, while the paratype came from Yangjian of Pan County, Guizhou Province. [1] An additional uncatalogued LPV specimen was also referred to S. yunguiensis. These specimens all came from the Lagerstätte discovered during a 2007 geological mapping project with a diverse record of marine life called Luoping Biota, that yielded, apart from Sinosaurosphargis, various invertebrates, fish, basal ichthyosaurs, Atopodentatus , the basal saurosphargid Largocephalosaurus , and several species of eosauropterygian, including both pachypleurosaurs and nothosaurids. [2]
Sinosaurosphargis was first described and named by Chun Li, Olivier Rieppel, Xiao-Chun Wu, Li-Jun Zhao and Li-Ting Wang in 2011 and the type species is Sinosaurosphargis yunguiensis. The generic name is derived from Greek sino, meaning "China", sauros, meaning "lizard", and sphargis, the old genus name for the leatherback turtle, (in reference to the many similarities Sinosaurosphargis shares with Saurosphargis , a taxon whose holotype was lost and was thus considered a mystery), following the pattern of Sinosauropteryx . Like the name of Saurosphargis itself, it also refers to Sinosaurosphargis' dorsal osteoderm "body armor" and broadened ribs forming a closed chest rib basket - traits which are seemingly transitional between turtles and other reptiles. The specific name yunguiensis is derived from the names of the neighboring Yunnan and Guizhou Provinces, where the fossils were found. [1]
Sinosaurosphargis was a small reptile, measuring 70–80 cm (2.3–2.6 ft) long without the missing tail. [3] It is a basal marine reptile, closely related to Saurosphargis from the Lower Muschelkalk of southwestern Poland and eastern Netherlands. Sinosaurosphargis possesses retracted external nostrils, and shows closed upper temporal fenestrae, and a ventrally open cheek. Its trunk region is short and rounded, and covered by a shield composed of small osteoderms, just like its neck and the proximal portions of all four limbs. Its back vertebrae show elongated, distally expanded transverse projections. The chest ribs are broadened and flat, contacting one another along their length, and forming a closed chest rib basket. [1] Li et al. (2014) found some of these traits to be shared with all saurosphargids, while others are exclusive to Saurosphargis and Sinosaurosphargis. These genera have broader and flatter rib baskets than basal saurosphargids like Largocephalosaurus , and also transverse projections of back vertebrae and proximal portions of chest ribs that are much more robust, being much wider than the spacing between them. [4]
The following cladogram is simplified after the phylogenetic analysis of Li et al. (2011) and shows the placement of Sinosaurosphargis within Sauria. Ichthyopterygia was removed from the tree, as it caused a less resolved topology. Saurosphargis was coded solely based on the holotype, as the Netherlands material remains unprepared. [1]
Li et al. (2014) presented an updated version of the analysis above, showing interrelationships of all known Saurosphargidae species. The removal / inclusion of Ichthyopterygia was found to affect the topology the most - switching the positions of the Eusaurosphargis+Helveticosaurus and Thalattosauriformes clades, and altering the positions of several taxa within Eosauropterygia, which are not shown. [4]
Dinocephalosaurus is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, D. orientalis, which was named by Chun Li in 2003. Unlike other long-necked protorosaurs, Dinocephalosaurus convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative Pectodens, which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, Dinocephalosaurus probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as suggested by the poorly-ossified and paddle-like limbs which would have prevented it from going ashore.
Thalattosauria is an extinct order of marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.
Miodentosaurus is a genus of thalattosaurian from the Late Triassic of China. It is one of several thalattosaurs found in the Xiaowa Formation, also known as the Wayao Member of the Falang Formation. The genus name "Miodentosaurus" translates to "Few toothed-lizard" while the species name "brevis" means "short", in reference to its short snout.
Chinchenia is an extinct genus of a basal pistosauroid known from the Middle Triassic of Guizhou Province, southwestern China. It contains a single species, Chinchenia sungi.
Helveticosaurus is an extinct genus of diapsid marine reptile known from the Middle Triassic of southern Switzerland. It contains a single species, Helveticosaurus zollingeri, known from the nearly complete holotype T 4352 collected at Cava Tre Fontane of Monte San Giorgio, an area well known for its rich record of marine life during the Middle Triassic.
The Guanling Formation is a Middle Triassic geologic formation in southwestern China.
Largocephalosaurus is an extinct genus of basal saurosphargid, a marine reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a type species, Largocephalosaurus polycarpon, and a second species L. qianensis.
Diandongosuchus is an extinct genus of archosauriform reptile, possibly a member of the Phytosauria, known from the Middle Triassic of China. The type species Diandongosuchus fuyuanensis was named in 2012 from the Zhuganpo Formation of Yunnan Province. It is a marine species that shows similarities with another Chinese Triassic species called Qianosuchus mixtus, although it has fewer adaptations toward marine life. It was originally classified as the basal-most member of the pseudosuchian clade Poposauroidea. However, a subsequent study conducted by Stocker et al. indicated it to be the basalmost known phytosaur instead.
Concavispina is an extinct genus of thalattosaur reptile from the early Late Triassic Xiaowa Formation of Guangling, Guizhou, southern China. It contains a single species, Concavispina biseridens. It is known only from the holotype ZMNH M8804, a nearly complete 364 cm long skeleton. Concavispina can be differentiated from other thalattosaurs by possessing two rows of blunt teeth on the anterior part of the maxilla and a V-shaped notch on the dorsal margin of each neural spine in the dorsal (back) vertebrae. Both its generic and specific names refer to these autapomorphies, as Concavispina means "concave spine" and biseridens means "two rows of teeth". It is thought to be most closely related to Xinpusaurus, as both taxa share three derived characters: a maxilla that is curved upward at its anterior end, a humerus that is wider near the shoulder than near the elbow, and the presence of less than five cervicals.
Atopodentatus is an extinct genus of basal sauropterygian known from the early Middle Triassic (Anisian) of Guanling Formation in Luoping County, Yunnan Province, southwestern China. It contains a single species, Atopodentatus unicus. It is thought to have lived between 247 and 240 million years ago, during the Middle Triassic period, about six million years after the Permian extinction. Atopodentatus was an herbivorous marine reptile, although marine reptiles are usually omnivores or carnivores.
Eretmorhipis is an extinct genus of hupehsuchian marine reptiles from the Early Triassic of China. It is currently known from two specimens that were discovered in an exposure of the Jialingjiang Formation in Yuan'an County, Hubei, and referred to the newly named species Eretmorhipis carrolldongi in 2015.
Sanchiaosaurus is an extinct genus of a basal nothosauroid known from the Middle Triassic of Guizhou Province, southwestern China. It contains a single species, Sanchiaosaurus dengi.
Saurosphargidae is an extinct family of marine reptiles known from the Early Triassic and early Middle Triassic of Europe and China.
Saurosphargis is an extinct genus of a basal marine reptile, saurosphargid, known from the Middle Triassic of southwestern Poland and eastern Netherlands. It contains a single species, Saurosphargis volzi.
Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi. It was a small reptile, measuring 20 cm (7.9 in) long.
Helveticosauridae is an extinct family of basal marine reptiles known from the Middle Triassic of southern Switzerland and northern Italy.
Pectodens is an extinct genus of archosauromorph reptile which lived during the Middle Triassic in China. The type and only species of the genus is P. zhenyuensis, named by Chun Li and colleagues in 2017. It was a member of the Archosauromorpha, specifically part of the unnatural grouping Protorosauria. However, an unusual combination of traits similar and dissimilar to other protorosaurs initially led to confusion over its evolutionary relationships. In 2021, it was placed in a newly-established group, Dinocephalosauridae, along with its closest relative Dinocephalosaurus.
The Xiaowa Formation is a Carnian-age geological formation found in southern China. It is a sequence of limestone and marls from the Carnian stage of the Triassic. Its lower section was previously known as the Wayao Formation or Wayao Member of the Falang Formation. In 2002, the Wayao Member was renamed and raised to the Xiaowa Formation to prevent confusion with an Eocene unit of the same name. Crinoids and marine reptiles are abundant in the Xiaowa Formation, forming a lagerstätte known as the Guanling biota. Ammonoids and conodonts found in the formation constrain its age to the early Carnian. Reptiles of the Guanling biota include ichthyosaurs, thalattosaurs, placodonts, and Odontochelys. Sedimentary events within this formation have been tied to the Carnian Pluvial Event.
The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.
Luopingosaurus is an extinct genus of pachypleurosaurid sauropterygian from the Middle Triassic Guanling Formation of Yunnan Province, China. The genus contains a single species, L. imparilis, known from a well-preserved, nearly complete skeleton.
{{cite journal}}
: CS1 maint: multiple names: authors list (link)