| Leptocleidus Temporal range: Early Cretaceous, | |
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| Life restoration of L. capensis | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Superorder: | † Sauropterygia |
| Order: | † Plesiosauria |
| Family: | † Leptocleididae |
| Genus: | † Leptocleidus Andrews, 1922 |
| Species | |
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| Synonyms | |
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Leptocleidus is an extinct genus of plesiosaur, [1] belonging to the family Leptocleididae. [2] It was a small plesiosaur, measuring only up to 3 m (9.8 ft). [3]
In short, the term Leptocleidus means "slender clavicle". It comes from a merge of the Greek words λεπτοσ, meaning "slender" and κλειδ (also spelled κλεισ) meaning clavicle.
Leptocleidus is known from the following sediments:
A specimen from the Vectis Formation (lower Aptian age), Isle of Wight, found in 1995 and seen as a "Leptocleidus sp.", was named as a separate genus Vectocleidus in 2012.
With large clavicles and interclavicle and small scapulae, Leptocleidus resembled the Early Jurassic Rhomaleosaurus and members of the Cretaceous family, Polycotylidae. The animal had 21 teeth on either side of its maxilla and approximately 35 teeth on each side of the mandible. The Leptocleidus' triangle-shaped skull had a crest running from a ridge on the end of the nose to the nasal region. Differing from pliosaurids, Leptocleidus had single-headed cervical ribs and a deep depression in the centra of the neck vertebrae. [3]
Leptocleidus, unlike many plesiosaurs, lived in shallow lagoons and likely visited brackish and fresh water systems (such as the mouths of large rivers). This led Arthur Richard Ivor Cruickshank to infer that this movement to fresh water was an attempt to flee larger plesiosaurs and pliosaurs. Most species are known from The British Isles but L. capensis was discovered in Cape Province, South Africa. [3]
Cladogram based on Ketchum and Benson (2011): [4]
| Leptocleididae |
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Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled plesiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae.
Rhomaleosaurus is an extinct genus of Early Jurassic rhomaleosaurid pliosauroid known from Northamptonshire and from Yorkshire of the United Kingdom. It was first named by Harry Seeley in 1874 and the type species is Rhomaleosaurus cramptoni. It was one of the earliest large marine reptile predators which hunted in the seas of Mesozoic era, measuring about 7 metres (23 ft) long. Like other pliosaurs, Rhomaleosaurus fed on ichthyosaurs, ammonites and other plesiosaurs.
Microcleidus is an extinct genus of sauropterygian reptile belonging to the Plesiosauroidea. The species has 40 neck vertebrae and a short tail of 28 vertebrae. Fossils of the genus have been found in France, the Posidonia Shale in Germany and Luxembourg, and the Alum Shale Formation of England.
Umoonasaurus is an extinct genus of plesiosaur belonging to the family Leptocleididae. This genus lived approximately 115 million years ago during the Early Cretaceous period, in shallow seas covering parts of what is now Australia. It was a relatively small animal around 2.5 metres (8.2 ft) long. An identifying trait of Umoonasaurus is three crest-ridges on its skull.
Plesiopleurodon is an extinct genus of Mesozoic marine reptiles, belonging to Sauropterygia, known from the Late Cretaceous of North America. It was named by Kenneth Carpenter based on a complete skull with a mandible, cervical vertebra, and a coracoid. In naming the specimen, Carpenter noted "Of all known pliosaurs, Plesiopleurodon wellesi most closely resembles Liopleurodon ferox from the Oxfordian of Europe, hence the generic reference." It was initially described as a pliosaur due to it short neck, a common trait of the family, although it is in the order Plesiosauria. However, later exploration into the relationships of both orders indicate that not all pliosaurs have short necks and not all plesiosaurs have long necks.
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874.
Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the Maastrichtian.
Brachauchenius is an extinct genus of pliosaurid that lived in North America and Morocco during the Late Cretaceous.
Nichollssaura is an extinct genus of leptocleidid plesiosaur from the Early Cretaceous Boreal Sea of North America. The type species is N. borealis, found in the early Albian age Clearwater Formation near Fort McMurray, Alberta, Canada.
Simolestes is an extinct pliosaurid genus that lived in the Middle to Late Jurassic. The type specimen, BMNH R. 3319 is an almost complete but crushed skeleton diagnostic to Simolestes vorax, dating back to the Callovian of the Oxford Clay formation, England. The genus might also be known from the Tithonian Bhuj Formation of India (S.indicus), however the referral of this species to Simolestes is dubious. S.keileni from France was moved to the new genus Lorrainosaurus in 2023.
Leptocleididae is a family of small-sized plesiosaurs that lived during the Early Cretaceous period. They had small bodies with small heads and short necks. Leptocleidus and Umoonasaurus had round bodies and triangle-shaped heads. Leptocleidids have been found in what were shallow nearshore, freshwater and brackish habitats. Hilary F. Ketchum and Roger B. J. Benson (2010), transferred Brancasaurus, Kaiwhekea, Nichollssaura and Thililua to this family. However, Ketchum and Benson (2011) reassigned Kaiwhekea and Thililua to their original positions, as an elasmosaurid and a polycotylid, respectively.
Hauffiosaurus is an extinct genus of Early Jurassic pliosaurid plesiosaur known from Holzmaden of Germany and from Yorkshire of the United Kingdom. It was first named by Frank Robin O’Keefe in 2001 and the type species is Hauffiosaurus zanoni. In 2011, two additional species were assigned to this genus: H. longirostris and H. tomistomimus.
Eurycleidus is an extinct genus of large-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, E. arcuatus. Like other plesiosaurs, Eurycleidus probably lived on a diet of fish, using its sharp needle-like teeth to catch prey. Its shoulder bones were fairly large, indicating a powerful forward stroke for fast swimming.
Rhomaleosauridae is a family of plesiosaurs from the Earliest Jurassic to the latest Middle Jurassic of Europe, North America, South America and possibly Asia. Most rhomaleosaurids are known from England, many specifically from lower Blue Lias deposits that date back to the earliest Jurassic, just at the boundary with the Triassic. In fact, to date only two undisputed rhomaleosaurids were named from outside Europe - the closely related Borealonectes russelli and Maresaurus coccai from Canada and Argentina, respectively. These two species are also the only Middle Jurassic representatives of the family. Rhomaleosauridae was formally named by Kuhn in 1961, originally proposed to include Rhomaleosaurus cramptoni and its relatives, which have short necks and large heads relatively to plesiosauroids like Elasmosaurus and Plesiosaurus, but longer necks and smaller heads relatively to advanced pliosaurids like Pliosaurus and Kronosaurus.
Microcleididae is an extinct family of basal plesiosauroid plesiosaurs from the Early Jurassic of France, Germany, Portugal and the United Kingdom. Currently, the oldest and the most known microcleidid is Eretmosaurus from the middle Sinemurian of the United Kingdom. Microcleididae was formally named and described by Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller in 2012.
Hastanectes is an extinct genus of a plesiosaurian with possible pliosaurid affinities known from the Early Cretaceous Wadhurst Clay Formation of the United Kingdom. It contains a single species, Hastanectes valdensis, which was originally thought to be a species of Cimoliasaurus.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Megacephalosaurus is an extinct genus of short-necked pliosaur that inhabited the Western Interior Seaway of North America about 94 to 93 million years ago during the Turonian stage of the Late Cretaceous, containing the single species M. eulerti. It is named after its large head, which is the largest of any plesiosaur in the continent and measures up to 1.75 meters (5.7 ft) in length. Megacephalosaurus was one of the largest marine reptiles of its time with an estimated length of 6–9 meters (20–30 ft). Its long snout and consistently sized teeth suggest that it preferred a diet of smaller-sized prey.
