Brachauchenius

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Brachauchenius
Temporal range: Late Cretaceous, 100–90  Ma
USNM 4989 - Brachauchenius lucasi holotype (in original position).png
Holotype specimen of B. lucasi (USNM 4989)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Sauropterygia
Order: Plesiosauria
Family: Pliosauridae
Subfamily: Brachaucheninae
Genus: Brachauchenius
Williston, 1903 [1]
Type species
Brachauchenius lucasi
Williston, 1903 [1]

Brachauchenius (meaning 'short neck') is an extinct genus of pliosaurid that lived in North America (United States) and Morocco [2] during the Late Cretaceous.

Contents

History

The second known skull of B. lucasi (USNM 2361) USNM 2361 - Brachauchenius lucasi skull.png
The second known skull of B. lucasi (USNM 2361)

The type species, Brachauchenius lucasi, lived in the Western Inland Sea of North America around 100-90 million years ago, from the Cenomonian to the Turonian of the Cretaceous. An older specimen from the Barremian of Colombia was considered as part of this genus, representing the first re-occurrence of the non-rhomaleosaurid pliosaurs after a Berriasian-Hauterivian hiatus. [3] However, subsequent analysis of this Colombian specimen shows that it was enough distinctive to warrant a new genus and species, named as Stenorhynchosaurus munozi . [4]

The first known (type) specimen (USNM 4989) was collected by Charles Hazelius Sternberg from Ottawa County, Kansas in 1884. [5] [6] It had a skull length of up to 1 m (3.3 ft). [7] The species was named by Samuel W. Williston. [1] [8] The individual to which this specimen belongs would have measured 4–5 m (13–16 ft) long. [7] Brachauchenius represents the last known occurrence of a pliosaur in North America.

A larger specimen (FHSM VP-321 - skull length 170 cm) was collected by George Fryer Sternberg in 1952 from the Fairport Chalk of Russell County, Kansas, and later described by Carpenter. [9] Schumacher and Everhart (2005) reported on the age and locality of both Kansas specimens. [10] This specimen is now reassigned to its own genus and species, Megacephalosaurus eulerti . [7]

In 2013, Benson et al. referred a partial cranium from the Chalk Group of England (previously referred to Polyptychodon ) to Brachauchenius indet. [11]

In 2015, a pliosaur mandible from Turonian deposits near Goulmima, Morocco were referred to the species Brachauchenius lucasi [2]

Classification

Brachauchenius lucasi holotype skull in dorsal, palatal and lateral views Brachauchenius lucasi skull.jpg
Brachauchenius lucasi holotype skull in dorsal, palatal and lateral views

The cladogram below follows a 2011 analysis by paleontologists Hilary F. Ketchum and Roger B. J. Benson, and reduced to genera only. [12]

B. lucasi pursuing a hesperornithiform bird Brachauchenius lucasi2DB.jpg
B. lucasi pursuing a hesperornithiform bird
Pliosauroidea

See also

Related Research Articles

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Plesiosauroidea is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous periods. After their discovery, some plesiosauroids were said to have resembled "a snake threaded through the shell of a turtle", although they had no shell.

<span class="mw-page-title-main">Pliosauroidea</span> Extinct clade of reptiles

Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled plesiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.

<span class="mw-page-title-main">Plesiosaur</span> Order of reptiles (fossil)

The Plesiosauria or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia.

Liopleurodon is an extinct genus of large, carnivorous marine reptile belonging to the Thalassophonea, a clade of short-necked pliosaurid plesiosaurs. Liopleurodon lived from the Callovian Stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic Period. It was the apex predator of the Middle to Late Jurassic seas that covered Europe. The largest species, L. ferox, is estimated to have grown to 6.4 metres (21 ft) in length based on a large skull.

<span class="mw-page-title-main">Elasmosauridae</span> Extinct family of reptiles

Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae.

<i>Plesiopleurodon</i> Extinct genus of reptiles

Plesiopleurodon is an extinct genus of Mesozoic marine reptiles, belonging to Sauropterygia, known from the Late Cretaceous of North America. It was named by Kenneth Carpenter based on a complete skull with a mandible, cervical vertebra, and a coracoid. In naming the specimen, Carpenter noted "Of all known pliosaurs, Plesiopleurodon wellesi most closely resembles Liopleurodon ferox from the Oxfordian of Europe, hence the generic reference." It was initially described as a pliosaur due to it short neck, a common trait of the family, although it is in the order Plesiosauria. However, later exploration into the relationships of both orders indicate that not all pliosaurs have short necks and not all plesiosaurs have long necks.

<span class="mw-page-title-main">Pliosauridae</span> Extinct family of reptiles

Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874.

<span class="mw-page-title-main">Polycotylidae</span> Extinct family of reptiles

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<i>Dolichorhynchops</i> Extinct genus of reptiles

Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species D. osborni and D. herschelensis, with two previous species having been assigned to new genera. Definitive specimens of D. osborni have been found in the late Coniacian to early Campanian rocks, while those of D. herschelensis have been found in the late Campanian to early Maastrichtian rocks. Dolichorhynchops was a prehistoric marine reptile measuring around 3 metres (9.8 ft) long. Its Greek generic name means "long-nosed face".

<i>Leptocleidus</i> Extinct genus of reptiles

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<i>Nichollssaura</i> Extinct genus of reptiles

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<i>Simolestes</i> Extinct genus of reptiles

Simolestes is an extinct pliosaurid genus that lived in the Middle to Late Jurassic. The type specimen, NHMUK PV R 3319 is an almost complete but crushed skeleton diagnostic to Simolestes vorax, dating back to the Callovian of the Oxford Clay formation, England. The genus might also be known from the Tithonian Bhuj Formation of India (S.indicus), however the referral of this species to Simolestes is dubious. S.keileni from France was moved to the new genus Lorrainosaurus in 2023.

<span class="mw-page-title-main">Leptocleididae</span> Extinct family of reptiles

Leptocleididae is a family of small-sized plesiosaurs that lived during the Early Cretaceous period. They had small bodies with small heads and short necks. Leptocleidus and Umoonasaurus had round bodies and triangle-shaped heads. Leptocleidids have been found in what were shallow nearshore, freshwater and brackish habitats. Hilary F. Ketchum and Roger B. J. Benson (2010), transferred Brancasaurus, Kaiwhekea, Nichollssaura and Thililua to this family. However, Ketchum and Benson (2011) reassigned Kaiwhekea and Thililua to their original positions, as an elasmosaurid and a polycotylid, respectively.

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<span class="mw-page-title-main">Timeline of plesiosaur research</span>

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<i>Megacephalosaurus</i> Extinct genus of reptiles

Megacephalosaurus is an extinct genus of short-necked pliosaur that inhabited the Western Interior Seaway of North America about 94 to 93 million years ago during the Turonian stage of the Late Cretaceous, containing the single species M. eulerti. It is named after its large head, which is the largest of any plesiosaur in the continent and measures up to 1.75 meters (5.7 ft) in length. Megacephalosaurus was one of the largest marine reptiles of its time with an estimated length of 6–9 meters (20–30 ft). Its long snout and consistently sized teeth suggest that it preferred a diet of smaller-sized prey.

<i>Stenorhynchosaurus</i> Extinct genus of reptiles

Stenorhynchosaurus is an extinct genus of pliosaurid plesiosaurs which lived in the Early Cretaceous of South America. The type species and only known is Stenorhynchosaurus munozi.

Luskhan is an extinct genus of brachauchenine pliosaur from the Cretaceous of Russia. The type and only species is Luskhan itilensis, named by Valentin Fischer and colleagues in 2017 from a well-preserved and nearly complete skeleton. As an early-diverging brachauchenine, Luskhan consequently exhibits an intermediate combination of traits seen in more basal and more derived pliosaurs. However, Luskhan departs significantly from other pliosaurs in that it exhibits a lack of adaptations in its skull to feeding on large prey; its slender snout, small teeth, and short tooth rows instead indicate a skull adapted for feeding on small, soft prey. With these features, it is the pliosaur that approaches closest to the distantly-related piscivorous polycotylids, having convergently evolved these traits more than 10 million years apart.

<i>Sachicasaurus</i> Extinct genus of pliosaurid

Sachicasaurus is an extinct genus of brachauchenine pliosaurid known from the Barremian of the Paja Formation, Altiplano Cundiboyacense in the Colombian Eastern Ranges of the Andes. The type species is S. vitae.

References

  1. 1 2 3 Williston SW. 1903. North American plesiosaurs. Field Columbian Museum, Pub. 73, Geological Series2 (1):1-79. (29 plates)
  2. 1 2 ANGST, D.; BARDET, N. (2015-08-05). "A new record of the pliosaurBrachauchenius lucasiWilliston, 1903 (Reptilia: Sauropterygia) of Turonian (Late Cretaceous) age, Morocco". Geological Magazine. 153 (3): 449–459. doi:10.1017/s0016756815000321. ISSN   0016-7568. S2CID   128840902.
  3. Hampe O. 2005. Considerations on a Brachauchenius skeleton (Pliosauroidea) from the lower Paja Formation (late Barremian) of Villa de Leyva area (Colombia). Fossil Record - Mitteilungen aus dem Museum für Naturkunde in Berlin8 (1): 37-51.
  4. Páramo, María E.; Gómez-Pérez, Marcela; Noé, Leslie F.; Etayo, Fernando (2016-04-06). "Stenorhynchosaurus munozi, gen. et sp. nov. a new pliosaurid from the Upper Barremian (Lower Cretaceous) of Villa de Leiva, Colombia, South America". Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales. 40 (154): 84–103. doi: 10.18257/raccefyn.239 . ISSN   2382-4980.
  5. Everhart MJ. 2005. Oceans of Kansas - A Natural History of the Western Interior Sea. Indiana University Press, 320 pp. ISBN   0-253-34547-2.
  6. Everhart MJ. 2007. Historical note on the 1884 discovery of Brachauchenius lucasi (Plesiosauria; Pliosauridae) in Ottawa County, Kansas. Transactions of the Kansas Academy of Science110 (3/4): 255-258.
  7. 1 2 3 4 Schumacher, B. A.; Carpenter, K.; Everhart, M. J. (2013). "A new Cretaceous Pliosaurid (Reptilia, Plesiosauria) from the Carlile Shale (middle Turonian) of Russell County, Kansas". Journal of Vertebrate Paleontology. 33 (3): 613. doi:10.1080/02724634.2013.722576. S2CID   130165209.
  8. Williston SW. 1907. The skull of Brachauchenius, with special observations on the relationships of the plesiosaurs. United States National Museum Proceedings32: 477-489. (pls. 34-37)
  9. Carpenter K. 1996. A Review of short-necked plesiosaurs from the Cretaceous of the western interior, North America. Neues Jahbruch für Geol. Palaont. Abh. (Stuttgart)201 (2): 259-287.
  10. Schumacher BA, Everhart MJ. 2005. A stratigraphic and taxonomic review of plesiosaurs from the old “Fort Benton Group” of central Kansas: A new assessment of old records. Paludicola5 (2): 33-54.
  11. Benson RBJ, Evans M, Smith AS, Sassoon J, Moore-Faye S, Ketchum HF, Forrest R. 2013. A giant pliosaurid skull from the Late Jurassic of England. PLOS ONE 8(5):e65989 DOI 10.1371/journal.pone.0065989.
  12. Hilary F. Ketchum; Roger B. J. Benson (2011). "A new pliosaurid (Sauropterygia, Plesiosauria) from the Oxford Clay Formation (Middle Jurassic, Callovian) of England: evidence for a gracile, longirostrine grade of Early-Middle Jurassic pliosaurids". Special Papers in Palaeontology. 86: 109–129.