Thalassiodracon Temporal range: | |
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Lectotype specimen NHMUK PV OR 2018, found in the Lower Lias strata, Street, Somerset, England. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Superorder: | † Sauropterygia |
Order: | † Plesiosauria |
Family: | † Pliosauridae |
Genus: | † Thalassiodracon Storrs & Taylor, 1996 |
Species: | †T. hawkinsii |
Binomial name | |
†Thalassiodracon hawkinsii (Owen, 1838) | |
Synonyms | |
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Thalassiodracon (tha-LAS-ee-o-DRAY-kon) is an extinct genus of plesiosauroid from the Pliosauridae that was alive during the Late Triassic-Early Jurassic (Rhaetian-Hettangian) and is known exclusively from the Lower Lias of England. [1] The type and only species, is Thalassiodracon ( Plesiosaurus ) hawkinsii (Owen, 1838). [2] [3]
Thalassiodracon hawkinsii is known from a number of complete skeletons (lectotype: NHMUK PV OR 2018) acquired by the fossil collector Thomas Hawkins in Somerset, England during the early 1830s, before 1834. Hawkins, an eccentric Pre-Adamite who had his fossils heavily restored and illustrated by distinguished artists in expensive editions to propagate his ideas, named these Plesiosaurus triotarsostinus in 1834 and Hezatarostinus in 1840 but these names are generally disregarded. [3] [4] [5] In Memoirs of Icthyosaurii and Plesiosaurii (1835) and The Book of the Great Sea Dragons (1840), Hawkins published his own illustrations after reconstructing the fossils he had obtained. [6] Some of Hawkins original notes are being stored at the Natural History Museum in London. [7] It was named as Plesiosaurus Hawkinsii in 1838 by Richard Owen [2] and it was made the type species of the genus Thalassiodracon in 1996 by Storrs & Taylor. [1]
The original fossil, designated as NHMUK PV OR 2018 was bought by Hawkins in the 1830s, and is currently on display at the Natural History Museum in London, England. A cast of the holotype fossil was created in 1862 by Henry Augustus Ward. This replica was catalogued as LDUCZ-X227 by Ray Lankester and is also located in London, at the Grant Museum of Zoology and Comparative Anatomy. Another cast of Thalassiodracon hawkinsi is being displayed at the Geological Museum of Trinity College in Dublin, Ireland. [8] Three fossils found in Somerset, England originally identified as Thalassiodracon were redescribed as another small plesiosaur, Stratesaurus taylori. [9] According to the Global Biodiversity Information Facility, there is a total of 31 fossil occurrences of Thalassiodracon hawkinsii, all located in the United Kingdom. [10]
The genus name Thalassiodracon means "sea dragon", while the specific name hawkinsii honours Hawkins.
Thalassiodracon was a small plesiosaur in the Late Triassic (Rhaetian) to the Early Jurassic (Hettangian) of Europe (age range: 201.6 to 196.5 million years ago). [3] It measured approximately 1.5 m (4.9 ft) long. [11] It has a skull length of 17 cm (6.7 in), neck length of 75.5 cm (2 ft 5.7 in), and trunk length of 68 cm (2 ft 3 in). [12]
Plesiosaurs lived in both saltwater and freshwater areas, with specific adaptations to help them thrive, with Thalassiodracon hawkinsi being no exception. The neck of Thalassiodracon was slightly shorter than that of subsequent plesiosaurs. [13] The cervical vertebrae were between 27 and 31, while those of Plesiosaurus were 35 - 37. Moreover, the skull was unusually short and equipped with long teeth. The orbits were very large, with sclerotic rings. The orbits of the skull were enlarged, containing likewise large, flat sclerotic rings. These rings are theorized to help the eyes of the creature to be well adjusted for a low-light environment found in its habitat. Other features like elongated staples and fused exoccipital bones seem to suggest adapted underwater hearing. [14] Like some modern reptiles, the teeth inside the skull had a constant replacement cycle as they wore out during the animal's lifetime. The cycle appears to have occurred in a wave shaped pattern, with cycle repeating every third tooth. [15]
A computed tomography of an exceptionally preserved skull, and examination of other specimens (Benson et al., 2011 [16] ) yields new anatomical data. Thalassiodracon had a dorsomedian ridge on the premaxilla, a squamosal bulb, four premaxillary teeth, and a heterodont maxillary dentition. [1] [17]
Like all plesiosaurs, this animal had legs like paddles for swimming in the sea. But, separating them from earlier ancestors, Thalassiodracon and following members also had shortened tails. [18] In adulthood, the forelimbs of Thalassiodracon hawkinsii are shorter than its hindlimbs. [19] Plesiosaurs with this characteristic are speculated to be pursuit predators, using speed and strength to hunt down their prey. In general, plesiosaurs had three primary ways to traverse through the water with their flippers: rowing, flying, and rowing flight. Unlike the reptiles of today, Thalassiodracon hawkinsii and other plesiosaurs were warm-blooded with a high metabolism, enabling them to have an active lifestyle. [18]
All plesiosaurs, including Thalassiodracon were faunivorous, but widely ranged in their diets. Animals such as ammonites, cephalopods, and other invertebrates were often found in plesiosaur remains. [20] With the bones of dinosaurs and pterosaurs being found in plesiosaur remains, these animals are also possibilities for their diets. [20]
Based off of other plesiosaurs, Thalassiodracon hawkinsii was viviparous, meaning it gave birth to live young. [21]
Thalassiodracon was originally placed in the genus Plesiosaurus . It has been classified in a new genus Thalassiodracon many years later following an examination of a skull. This animal is considered one of the oldest representatives of the group of plesiosaurs, perhaps ancestral to the superfamily of Pliosauroidea (short-necked plesiosaurs). [22]
The following cladogram follows an analysis by Ketchum & Benson, 2011. [16]
Plesiosaurus is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias of England. It is distinguishable by its small head, long and slender neck, broad turtle-like body, a short tail, and two pairs of large, elongated paddles. It lends its name to the order Plesiosauria, of which it is an early, but fairly typical member. It contains only one species, the type, Plesiosaurus dolichodeirus. Other species once assigned to this genus, including P. brachypterygius, P. guilielmiimperatoris, and P. tournemirensis have been reassigned to new genera, such as Hydrorion, Seeleyosaurus and Occitanosaurus.
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled plesiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
The Plesiosauria or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia.
Pistosaurus is an extinct genus of aquatic sauropterygian reptile closely related to plesiosaurs. Fossils have been found in France and Germany, and date to the Middle Triassic. It contains a single species, Pistosaurus longaevus. Pistosaurus is known as the oldest "subaquatic flying" reptile on earth.
Peloneustes is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of Plesiosaurus by palaeontologist Harry Govier Seeley in 1869, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to Peloneustes, P. philarchus is currently the only one still considered valid, with the others moved to different genera, considered nomina dubia, or synonymised with P. philarchus. Some of the material formerly assigned to P. evansi have since been reassigned to "Pliosaurus" andrewsi. Peloneustes is known from many specimens, including some very complete material.
Rhomaleosaurus is an extinct genus of Early Jurassic rhomaleosaurid pliosauroid known from Northamptonshire and from Yorkshire of the United Kingdom. It was first named by Harry Seeley in 1874 and the type species is Rhomaleosaurus cramptoni. It was one of the earliest large marine reptile predators which hunted in the seas of Mesozoic era, measuring about 7 metres (23 ft) long. Like other pliosaurs, Rhomaleosaurus fed on ichthyosaurs, ammonites and other plesiosaurs.
Colymbosaurus is a genus of cryptoclidid plesiosaur from the Late Jurassic (Callovian-Tithonian) of the UK and Svalbard, Norway. There are two currently recognized species, C. megadeirus and C. svalbardensis.
Archaeonectrus is an extinct genus of pliosaur from the Early Jurassic (Sinemurian) of what is now southeastern England. The type species is Archaeonectrusrostratus, first named by Sir Richard Owen in 1865, which was moved to its own genus by N.I. Novozhilov in 1964. It was a relatively small plesiosaur, measuring 3.4–3.67 m (11.2–12.0 ft) long.
The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassic times, between 195 and 200 million years ago. The Blue Lias is famous for its fossils, especially ammonites.
Hauffiosaurus is an extinct genus of Early Jurassic pliosaurid plesiosaur known from Holzmaden of Germany and from Yorkshire of the United Kingdom. It was first named by Frank Robin O’Keefe in 2001 and the type species is Hauffiosaurus zanoni. In 2011, two additional species were assigned to this genus: H. longirostris and H. tomistomimus.
Eurycleidus is an extinct genus of large-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, E. arcuatus. Like other plesiosaurs, Eurycleidus probably lived on a diet of fish, using its sharp needle-like teeth to catch prey. Its shoulder bones were fairly large, indicating a powerful forward stroke for fast swimming.
Eretmosaurus is an extinct genus of plesiosaur from the Early Jurassic Blue Lias of England. Only the type species is known, which is E. rugosus.
Rhomaleosauridae is a family of plesiosaurs from the Earliest Jurassic to the latest Middle Jurassic of Europe, North America, South America and possibly Asia. Most rhomaleosaurids are known from England, many specifically from lower Blue Lias deposits that date back to the earliest Jurassic, just at the boundary with the Triassic. In fact, to date only two undisputed rhomaleosaurids were named from outside Europe - the closely related Borealonectes russelli and Maresaurus coccai from Canada and Argentina, respectively. These two species are also the only Middle Jurassic representatives of the family. Rhomaleosauridae was formally named by Kuhn in 1961, originally proposed to include Rhomaleosaurus cramptoni and its relatives, which have short necks and large heads relatively to plesiosauroids like Elasmosaurus and Plesiosaurus, but longer necks and smaller heads relatively to advanced pliosaurids like Pliosaurus and Kronosaurus.
Avalonnectes is an extinct genus of small-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, A. arturi.
Stratesaurus is an extinct genus of small-bodied rhomaleosaurid plesiosaur known from the Early Jurassic period of the United Kingdom. It contains a single species, S. taylori. It was a small plesiosaur, with a skull length of 18 cm (7.1 in) and a body length of 2 m (6.6 ft).
Eoplesiosaurus is an extinct genus of basal plesiosauroid known from the Early Jurassic period of the United Kingdom. It contains a single species, E. antiquior.
Anningasaura is an extinct genus of basal plesiosaur. It is known from a single type species, A. lymense, discovered in Early Jurassic rocks of Lyme Regis in the United Kingdom.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Atychodracon is an extinct genus of rhomaleosaurid plesiosaurian known from the Late Triassic - Early Jurassic boundary of England. It contains a single species, Atychodracon megacephalus, named in 1846 originally as a species of Plesiosaurus. The holotype of "P." megacephalus was destroyed during a World War II air raid in 1940 and was later replaced with a neotype. The species had a very unstable taxonomic history, being referred to four different genera by various authors until a new genus name was created for it in 2015. Apart from the destroyed holotype and its three partial casts, a neotype and two additional individuals are currently referred to Atychodracon megacephalus, making it a relatively well represented rhomaleosaurid.