Scalamagnus

Last updated

Scalamagnus
Temporal range: Late Cretaceous (early Turonian), 93.5–92.0  Ma
O
S
D
C
P
T
J
K
Pg
N
Scalamagnus tropicensis.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Sauropterygia
Order: Plesiosauria
Family: Polycotylidae
Subfamily: Polycotylinae
Genus: Scalamagnus
Clark, O’Keefe & Slack, 2023
Type species
Scalamagnus tropicensis
Schmeisser McKean, 2012
Synonyms

Scalamagnus is an extinct genus of polycotylid plesiosaur from the Late Cretaceous Tropic Shale Formation of the United States. The genus contains a single species S. tropicensis, known from a skull and two partial skeletons. Scalamagnus was historically considered to represent a species of the genus Dolichorhynchops before it was moved to its own genus. [1]

Contents

History

The taxon was first named as a species of Dolichorhynchops by Rebecca Schmeisser McKean in 2012. The specific name is derived from the name of the Tropic Shale, in which the two specimens were found. It is known from the holotype MNA  V10046, an almost complete, well-preserved 3.2 metres (10 ft) long skeleton including the most of the skull and from the referred specimen MNA V9431, fragmentary postcranial elements. It was collected by the Museum of Northern Arizona from a single locality within the Tropic Shale of Utah, dating to the early Turonian stage of the early Late Cretaceous, about 93.5-91 million years ago. D. tropicensis extends the known stratigraphic range for Dolichorhynchops back by approximately 7 million years. [2] Previously three additional polycotylid taxa, Eopolycotylus , Palmulasaurus and Trinacromerum , have been named from the same formation, two of which are currently endemic to the Tropic Shale. [3] A 2023 study assigns D. tropicensis to a new genus, Scalamagnus. [1]

Classification

Clark, O’Keefe & Slack (2023) recovered Scalamagnus as a polycotylid member of the plesiosaur clade Leptocleidia, as the sister taxon to the clade formed by Trinacromerum and the Dolichorhynchia within the Polycotylinae. The results of their phylogenetic analyses are shown in the cladogram below: [1]

Leptocleidia
Brancasaurus
Brancasaurus brancai.png
Vectocleidus
Vectocleidus pastorum.png
Leptocleididae
Leptocleidus capensis.png
Polycotylidae
Edgarosaurus
Edgarosaurus muddi.png
Palmulasaurus
Palmulasaurus quadratus.png
Pahasapasaurus
Pahasapasaurus haasi.png
Occultonectia
Sulcusuchus erraini.png
Manemergus
Manemergus anguirostris.png
Thililua
Thililua longicollis.png
Polycotylinae
Eopolycotylus
Eopolycotylus rankini.png
Polycotylus latipinnis
Polycotylus latipinnis.png

Scalamagnus

Trinacromerum bentonianum
Trinacromerum bentonianum.png
Dolichorhynchia
Dolichorhynchops osborni
Dolichorhynchops osborni.png

Dolichorhynchops herschelensis

Martinectes

ROM 29010 (Niobrara polycotyline) [4]

Unktaheela

Paleobiology

Life restoration of Scalamagnus being chased by Megacephalosaurus Megacephalosaurus eulerti.jpg
Life restoration of Scalamagnus being chased by Megacephalosaurus

The holotype is associated with 289 gastroliths, which is unusual in comparison to most polycotylid skeletons that generally lack gastroliths. Ranging from less than 0.1 grams to 18.5 grams, the total mass of the gastroliths was about 518 grams. About three-quarters of the stones weighed less than 2 grams, with the mean mass and median mass of the stones respectively estimated at 1.9 grams and 0.8 grams. The gastroliths had high mean value and variability in sphericity, suggesting that this individual was obtaining its stones from rivers located along the western side of the Western Interior Seaway. [5]

Related Research Articles

<span class="mw-page-title-main">Plesiosaur</span> Order of reptiles (fossil)

The Plesiosauria or Plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia.

<span class="mw-page-title-main">Elasmosauridae</span> Extinct family of reptiles

Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae. Their diet mainly consisted of crustaceans and molluscs.

<i>Plesiopleurodon</i> Extinct genus of reptiles

Plesiopleurodon is an extinct genus of Mesozoic marine reptiles, belonging to Sauropterygia, known from the Late Cretaceous of North America. It was named by Kenneth Carpenter based on a complete skull with a mandible, cervical vertebra, and a coracoid. In naming the specimen, Carpenter noted "Of all known pliosaurs, Plesiopleurodon wellesi most closely resembles Liopleurodon ferox from the Oxfordian of Europe, hence the generic reference." It was initially described as a pliosaur due to it short neck, a common trait of the family, although it is in the order Plesiosauria. However, later exploration into the relationships of both orders indicate that not all pliosaurs have short necks and not all plesiosaurs have long necks.

<span class="mw-page-title-main">Polycotylidae</span> Extinct family of reptiles

Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the Maastrichtian.

<i>Thililua</i> Extinct genus of reptiles

Thililua is a genus of polycotylid plesiosaur, containing one species, T. longicollis.

<i>Dolichorhynchops</i> Extinct genus of reptiles

Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species D. osborni and D. herschelensis, with two previous species having been assigned to new genera. Specimens of D. osborni have been found in the early Coniacian to early Campanian rocks, while those of D. herschelensis have been found in the late Campanian to early Maastrichtian rocks. Dolichorhynchops was a prehistoric marine reptile. Its Greek generic name means "long-nosed face". While typically measuring about 3 metres (9.8 ft) in length, the largest specimen of D. osborni is estimated to have a total body length more than approximately 4.3 metres (14 ft).

<span class="mw-page-title-main">Bearpaw Formation</span> Geologic formation in North America

The Bearpaw Formation, also called the Bearpaw Shale, is a geologic formation of Late Cretaceous (Campanian) age. It outcrops in the U.S. state of Montana, as well as the Canadian provinces of Alberta and Saskatchewan, and was named for the Bear Paw Mountains in Montana. It includes a wide range of marine fossils, as well as the remains of a few dinosaurs. It is known for its fossil ammonites, some of which are mined in Alberta to produce the organic gemstone ammolite.

Terminonatator is a genus of elasmosaurid plesiosaur from the Late Cretaceous of Saskatchewan, Canada. It is known from a skull and partial skeleton from a young adult, found in the Campanian-age Bearpaw Formation near Notukeu Creek in Ponteix. Terminonatator is currently one of the youngest plesiosaurs from the Western Interior Seaway.

<span class="mw-page-title-main">Pierre Shale</span> Geologic formation of the Upper Cretaceous from Pembina Valley in Canada to New Mexico, USA

The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico.

<i>Brancasaurus</i> Extinct genus of reptiles

Brancasaurus is a genus of plesiosaur which lived in a freshwater lake in the Early Cretaceous of what is now North Rhine-Westphalia, Germany. With a long neck possessing vertebrae bearing distinctively-shaped "shark fin"-shaped neural spines, and a relatively small and pointed head, Brancasaurus is superficially similar to Elasmosaurus, albeit smaller in size at 3.26 metres (10.7 ft) in length as a subadult.

<i>Polycotylus</i> Extinct genus of reptiles

Polycotylus is a genus of plesiosaur within the family Polycotylidae. The type species is P. latippinis and was named by American paleontologist Edward Drinker Cope in 1869. Eleven other species have been identified. The name means 'much-cupped vertebrae', referring to the shape of the vertebrae. It lived in the Western Interior Seaway of North America toward the end of the Cretaceous. One fossil preserves an adult with a single large fetus inside of it, indicating that Polycotylus gave live birth, an unusual adaptation among reptiles.

<i>Palmulasaurus</i> Extinct genus of reptiles

Palmulasaurus is a genus of polycotylid plesiosaur from the Turonian Tropic Shale of Utah. It was originally described as Palmula, but the name was occupied by a genus of Cretaceous foraminifer first described in 1833.

Palmula is an extinct genus of foraminifera which is known from a number of species found in rocks dating from near the beginning of the Jurassic to the end of the Cretaceous, in Africa, Asia, Europe, and New Zealand. A genus of polycotylid plesiosaur was named Palmula in 2007, but because the name was already in use, the plesiosaur was renamed, becoming Palmulasaurus.

<span class="mw-page-title-main">Timeline of plesiosaur research</span>

This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.

<i>Mauriciosaurus</i> Extinct genus of reptiles

Mauriciosaurus is a genus of polycotylid plesiosaur from the Late Cretaceous of Mexico. It contains a single species, M. fernandezi, described in 2017 by Eberhard Frey and colleagues from a single well-preserved juvenile specimen about 1.9 metres long. Morphologically, it is overall most similar to the polycotyline polycotylids Trinacromerum and Dolichorhynchops. However, several features separate Mauriciosaurus from all other polycotylids, warranting the naming of a new genus. These include the sophisticated pattern of ridges on the bottom of the parasphenoid bone on its palate; the narrow openings in the palate bordered by the pterygoid bones; the lack of perforations in the surface of the coracoid; and the highly unusual arrangement of gastralia, or belly ribs, which is only otherwise seen in the non-polycotylid Cryptoclidus.

<i>Serpentisuchops</i> Genus of polycotylid plesiosaurs

Serpentisuchops is a genus of polycotylid plesiosaur from the late Cretaceous Pierre Shale of Wyoming, United States. The genus contains a single species, S. pfisterae, known from a partial skeleton.

<i>Unktaheela</i> Genus of polycotylid plesiosaurs

Unktaheela is an extinct genus of polycotylid plesiosaur from the Late Cretaceous Sharon Springs Formation of the United States. The genus contains a single species, U. specta, known from two partial skeletons. Unktaheela represents the smallest known polycotylid.

<i>Martinectes</i> Genus of polycotylid plesiosaurs

Martinectes is an extinct genus of polycotylid plesiosaur from the Late Cretaceous Sharon Springs Formation of the United States. The genus contains a single species M. bonneri, known from multiple skeletons and skulls. Martinectes was historically considered to represent a species of the genus Trinacromerum and later Dolichorhynchops before it was moved to its own genus.

This list of fossil reptiles described in 2024 is a list of new taxa of fossil reptiles that were described during the year 2024, as well as other significant discoveries and events related to reptile paleontology that occurred in 2024.

References

  1. 1 2 3 Clark, Robert O.; O’Keefe, F. Robin; Slack, Sara E. (2023-12-24). "A new genus of small polycotylid plesiosaur from the Upper Cretaceous of the Western Interior Seaway and a clarification of the genus Dolichorhynchops". Cretaceous Research : 105812. doi:10.1016/j.cretres.2023.105812. ISSN   0195-6671.
  2. Schmeisser McKean, Rebecca (2012). "A new species of polycotylid plesiosaur (Reptilia: Sauropterygia) from the Lower Turonian of Utah: extending the stratigraphic range of Dolichorhynchops". Cretaceous Research. 34: 184–199. doi:10.1016/j.cretres.2011.10.017.
  3. Albright III, L. B.; Gillette, D. D.; Titus, A. L. (2007b). "Plesiosaurs from the Upper Cretaceous (Cenomanian-Turonian) Tropic Shale of southern Utah, part 2: polycotylidae" (PDF). Journal of Vertebrate Paleontology. 27 (1): 41–58. doi:10.1671/0272-4634(2007)27[41:PFTUCC]2.0.CO;2. S2CID   130268187. Archived from the original (PDF) on 2011-09-28.
  4. Sato, Tamaki; Wu, Xiao-Chun; Tirabasso, Alex; Bloskie, Paul (2011-03-17). "Braincase of a polycotylid plesiosaur (Reptilia: Sauropterygia) from the Upper Cretaceous of Manitoba, Canada". Journal of Vertebrate Paleontology . 31 (2): 313–329. doi:10.1080/02724634.2011.550358. ISSN   0272-4634.
  5. Schmeisser, R.L.; Gillette, D.D. (2009). "Unusual occurrence of gastroliths in a polycotylid plesiosaur from the Upper Cretaceous Tropic Shale, southern Utah". PALAIOS. 24 (7): 453–459. Bibcode:2009Palai..24..453S. doi:10.2110/palo.2008.p08-085r. S2CID   128969768.