Polycotylids Temporal range: Early - Late Cretaceous, | |
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Martinectes in the Rocky Mountain Dinosaur Resource Center | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Superorder: | † Sauropterygia |
Order: | † Plesiosauria |
Clade: | † Leptocleidia |
Family: | † Polycotylidae Williston, 1909 |
Genera | |
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Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the early Maastrichtian around 72 million years ago. [3] The possible latest surviving member Rarosaurus from the late Maastrichtian is more likely a crocodylomorph. [4]
With their short necks and large elongated heads, they resemble the pliosaurs, but closer phylogenetic studies indicate that they share many common features with the Leptocleididae and Elasmosauridae. They have been found worldwide, with specimens reported from New Zealand, Australia, Japan, Morocco, the US, Canada, Eastern Europe, and South America. [5]
Cladogram after Albright, Gillette and Titus (2007). [6]
Plesiosauria |
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Cladogram after Ketchum and Benson (2010). [8]
Below is a cladogram of polycotylid relationships from Ketchum & Benson, 2011. [9]
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled plesiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae.
Microcleidus is an extinct genus of sauropterygian reptile belonging to the Plesiosauroidea. The species has 40 neck vertebrae and a short tail of 28 vertebrae. Fossils of the genus have been found in France, the Posidonia Shale in Germany and Luxembourg, and the Alum Shale Formation of England.
Umoonasaurus is an extinct genus of plesiosaur belonging to the family Leptocleididae. This genus lived approximately 115 million years ago during the Early Cretaceous period, in shallow seas covering parts of what is now Australia. It was a relatively small animal around 2.5 metres (8.2 ft) long. An identifying trait of Umoonasaurus is three crest-ridges on its skull.
Plesiopleurodon is an extinct genus of Mesozoic marine reptiles, belonging to Sauropterygia, known from the Late Cretaceous of North America. It was named by Kenneth Carpenter based on a complete skull with a mandible, cervical vertebra, and a coracoid. In naming the specimen, Carpenter noted "Of all known pliosaurs, Plesiopleurodon wellesi most closely resembles Liopleurodon ferox from the Oxfordian of Europe, hence the generic reference." It was initially described as a pliosaur due to it short neck, a common trait of the family, although it is in the order Plesiosauria. However, later exploration into the relationships of both orders indicate that not all pliosaurs have short necks and not all plesiosaurs have long necks.
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874.
Cryptoclididae is a family of medium-sized plesiosaurs that existed from the Middle Jurassic to the Early Cretaceous. They had long necks, broad and short skulls and densely packed teeth. They fed on small soft-bodied preys such as small fish and crustaceans. The earliest members of the family appeared during the early Bajocian, and they represented the dominant group of long-necked plesiosaurs during the latter half of the Jurassic.
Thililua is a genus of polycotylid plesiosaur, containing one species, T. longicollis.
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species D. osborni and D. herschelensis, with two previous species having been assigned to new genera. Definitive specimens of D. osborni have been found in the late Coniacian to early Campanian rocks, while those of D. herschelensis have been found in the late Campanian to early Maastrichtian rocks. Dolichorhynchops was a prehistoric marine reptile measuring around 3 metres (9.8 ft) long. Its Greek generic name means "long-nosed face".
Brachauchenius is an extinct genus of pliosaurid that lived in North America and Morocco during the Late Cretaceous.
Leptocleidus is an extinct genus of plesiosaur, belonging to the family Leptocleididae. It was a small plesiosaur, measuring only up to 3 m (9.8 ft).
Nichollssaura is an extinct genus of leptocleidid plesiosaur from the Early Cretaceous Boreal Sea of North America. The type species is N. borealis, found in the early Albian age Clearwater Formation near Fort McMurray, Alberta, Canada.
Archaeonectrus is an extinct genus of pliosaur from the Early Jurassic (Sinemurian) of what is now southeastern England. The type species is Archaeonectrusrostratus, first named by Sir Richard Owen in 1865, which was moved to its own genus by N.I. Novozhilov in 1964. It was a relatively small plesiosaur, measuring 3.4–3.67 m (11.2–12.0 ft) long.
Polyptychodon is a genus of pliosaurid found in Middle-Late Cretaceous marine deposits in southern England, France and Argentina. It has been considered a nomen dubium in a 2016 review.
Leptocleididae is a family of small-sized plesiosaurs that lived during the Early Cretaceous period. They had small bodies with small heads and short necks. Leptocleidus and Umoonasaurus had round bodies and triangle-shaped heads. Leptocleidids have been found in what were shallow nearshore, freshwater and brackish habitats. Hilary F. Ketchum and Roger B. J. Benson (2010), transferred Brancasaurus, Kaiwhekea, Nichollssaura and Thililua to this family. However, Ketchum and Benson (2011) reassigned Kaiwhekea and Thililua to their original positions, as an elasmosaurid and a polycotylid, respectively.
Eurycleidus is an extinct genus of large-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, E. arcuatus. Like other plesiosaurs, Eurycleidus probably lived on a diet of fish, using its sharp needle-like teeth to catch prey. Its shoulder bones were fairly large, indicating a powerful forward stroke for fast swimming.
Rhomaleosauridae is a family of plesiosaurs from the Earliest Jurassic to the latest Middle Jurassic of Europe, North America, South America and possibly Asia. Most rhomaleosaurids are known from England, many specifically from lower Blue Lias deposits that date back to the earliest Jurassic, just at the boundary with the Triassic. In fact, to date only two undisputed rhomaleosaurids were named from outside Europe - the closely related Borealonectes russelli and Maresaurus coccai from Canada and Argentina, respectively. These two species are also the only Middle Jurassic representatives of the family. Rhomaleosauridae was formally named by Kuhn in 1961, originally proposed to include Rhomaleosaurus cramptoni and its relatives, which have short necks and large heads relatively to plesiosauroids like Elasmosaurus and Plesiosaurus, but longer necks and smaller heads relatively to advanced pliosaurids like Pliosaurus and Kronosaurus.
Hastanectes is an extinct genus of a plesiosaurian with possible pliosaurid affinities known from the Early Cretaceous Wadhurst Clay Formation of the United Kingdom. It contains a single species, Hastanectes valdensis, which was originally thought to be a species of Cimoliasaurus.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.