| Polyptychodon Temporal range: Late Cretaceous, | |
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| P. interruptus jaw with tooth | |
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| Genus: | †Polyptychodon |
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| †Polyptychodon interruptus Owen, 1841 [1] | |
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Polyptychodon (meaning 'many-folded tooth') is a genus of pliosaurid found in Middle-Late Cretaceous marine deposits in southern England, France and Argentina. It has been considered a nomen dubium in a 2016 review. [2]
The type species, P. interruptus is known from an isolated tooth from the Late Cretaceous Chalk Group of southern England. [1] Owen described a second nominal species of the genus, P. continuus, from an isolated tooth collected in the Hythe Formation of Maidstone, Kent. (The macronarian sauropod Dinodocus was mistakenly thought to be conspecific with P. continuus before it was correctly recognized as a dinosaur and not a plesiosaur.)
Numerous pliosaurid teeth and vertebrae from England and eastern France have been previously assigned to Polyptychodon, including isolated vertebrae from France which were misidentified as a sauropod. [3] Comparison between Albian-age isolated vertebrae from marine deposits in France and Kronosaurus suggested a size of approximately 7 metres (23 ft) for a Polyptychodon-like brachaucheniine pliosaurid. [3] However, a 2016 re-evaluation found Polyptychodon and its types species to be dubious, and that numerous remains from the Chalk Group in England that had been referred to the genus most likely represent different species of plesiosaurs, with some teeth possibly being referable to Polycotylidae. [2] Similar fossils of pliosaurs were found also in Czech Republic. [4]
The species Polyptychodon patagonicus (Ameghino, 1893), based on crocodile teeth discovered in Argentina, shares the same genus name. According to a 2010 study, P. patagonicus is a nomen vanum and a nomen dubium . [5]
Polyptychodon hudsoni (holotytpe, SMU 60313) was described from the Turonian-age Eagle Ford Formation of Dallas, Texas. [6] [7] It probably belongs to a different genus. [2] [8]
Elasmosaurus is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5 million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was sent to the American paleontologist Edward Drinker Cope, who named it E. platyurus in 1868. The generic name means "thin-plate reptile", and the specific name means "flat-tailed". Cope originally reconstructed the skeleton of Elasmosaurus with the skull at the end of the tail, an error which was made light of by the paleontologist Othniel Charles Marsh, and became part of their "Bone Wars" rivalry. Only one incomplete Elasmosaurus skeleton is definitely known, consisting of a fragmentary skull, the spine, and the pectoral and pelvic girdles, and a single species is recognized today; other species are now considered invalid or have been moved to other genera.
Kronosaurus is an extinct genus of large short-necked pliosaur that lived during the Aptian to Albian stages of the Early Cretaceous in what is now Australia. The first known specimen was received in 1899 and consists of a partially preserved mandibular symphysis, which was first thought to come from an ichthyosaur according to Charles De Vis. However, it was 1924 that Albert Heber Longman formally described this specimen as the holotype of an imposing pliosaurid, to which he gave the scientific name K. queenslandicus, which is still the only recognized species nowadays. The genus name, meaning "lizard of Kronos", refers to its large size and possible ferocity reminiscent of the Titan of the Greek mythology, while the species name alludes to Queensland, the Australian state of its discovery. In the early 1930s, the Harvard Museum of Comparative Zoology sent an organized expedition to Australia that recovered two specimens historically attributed to the taxon, including a well known skeleton that is now massively restored in plaster. Several attributed fossils were subsequently discovered, including two large, more or less partials skeletons. As the holotype specimen does not present diagnostics to concretely distinguish Kronosaurus from other pliosaurids, these same two skeletons are proposed as potential neotypes for future redescriptions. Two additional species were proposed, but these are now seen as unlikely or belonging to another genus.
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled plesiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
Liopleurodon is an extinct genus of carnivorous pliosaurid pliosaurs that lived from the Callovian stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic period. The type species is L. ferox, which is probably the only valid species. Some studies also include the second species L. pachydeirus, but this latter is considered as a probable junior synonym of L. ferox due to its lack of viable diagnosis. As the holotype specimen of L. ferox consists of a single tooth preserving questionable distinctive features, recent studies therefore recommend the necessary identification of a neotype in order to preserve the validity of the genus. Numerous fossil specimens attributed to Liopleurodon, even including numerous skeletons, have been discovered in Europe, Russia, and Mexico. Other additional species were even proposed, but these are currently seen as coming from other pliosaurid genera.
Dinodocus is a genus of sauropod dinosaur, named by Richard Owen in 1884. The name is now usually considered a nomen dubium. The only species, D. mackesoni, a name given to some fossil bones from the Lower Greensand Group of Hythe, Kent, England, were formerly placed in the genus Pelorosaurus, but a review by Upchurch et al. (2004) concluded that Dinodocus is a nomen dubium.
Pukyongosaurus is a genus of titanosauriform dinosaur that lived in South Korea during the Early Cretaceous Period. It may have been closely related to Euhelopus, and is known from a series of vertebrae in the neck and back. The characteristics that were originally used to distinguish this genus have been criticized as being either widespread or too poorly preserved to evaluate, rendering the genus an indeterminate nomen dubium among titanosauriforms. The 2022 study noted that Pukyongosaurus is probably a somphospondylan.
Macrurosaurus is the name given to a genus of dinosaur from the Early Cretaceous. It was a titanosauriform which lived in what is now England. The type species, M. semnus, was named in 1876. A second species, M. platypus, may also exist.
Peloneustes is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of Plesiosaurus by palaeontologist Harry Govier Seeley in 1869, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to Peloneustes, P. philarchus is currently the only one still considered valid, with the others moved to different genera, considered nomina dubia, or synonymised with P. philarchus. Some of the material formerly assigned to P. evansi have since been reassigned to "Pliosaurus" andrewsi. Peloneustes is known from many specimens, including some very complete material.
Umoonasaurus is an extinct genus of plesiosaur belonging to the family Leptocleididae. This genus lived approximately 115 million years ago during the Early Cretaceous period, in shallow seas covering parts of what is now Australia. It was a relatively small animal around 2.5 metres (8.2 ft) long. An identifying trait of Umoonasaurus is three crest-ridges on its skull.
Woolungasaurus is a dubious genus of plesiosaur, an extinct marine reptile, belonging to the Elasmosauridae.
Brachauchenius is an extinct genus of pliosaurs that lived during the Late Cretaceous in what are now North America and North Africa. Only one species is known, B. lucasi, initially described by Samuel Wendell Williston in 1903 from a partial fossil skeleton discovered in a quarry in Kansas, United States. Many other fossil specimens attributed to the species were subsequently discovered, including an individual from Morocco whose presence was made official in 2016. Many contemporary pliosaur specimens were formerly attributed to Brachauchenius, but have since been reidentified as belonging to other genera or are recognized as indeterminate.
Polycotylus is a genus of plesiosaur within the family Polycotylidae. The type species is P. latippinis and was named by American paleontologist Edward Drinker Cope in 1869. Eleven other species have been identified. The name means 'much-cupped vertebrae', referring to the shape of the vertebrae. It lived in the Western Interior Seaway of North America toward the end of the Cretaceous. One fossil preserves an adult with a single large fetus inside of it, indicating that Polycotylus gave live birth, an unusual adaptation among reptiles.
Cardabiodon is an extinct genus of large mackerel shark that lived about 95 to 91 million years ago (Ma) during the Cenomanian to Turonian of the Late Cretaceous. It is a member of the Cardabiodontidae, a family unique among mackerel sharks due to differing dental structures, and contains the two species C. ricki and C. venator. Cardabiodon fossils have been found in Australia, North America, England, and Kazakhstan. It was likely an antitropical shark that inhabited temperate neritic and offshore oceans between 40° and 60° paleolatitude, similar to the modern porbeagle shark.
Rugocaudia is a potentially dubious genus of basal titanosauriform sauropod dinosaur known from the Early Cretaceous of Montana, United States.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Megacephalosaurus is an extinct genus of short-necked pliosaur that inhabited the Western Interior Seaway of North America about 94 to 93 million years ago during the Turonian stage of the Late Cretaceous, containing the single species M. eulerti. It is named after its large head, which is the largest of any plesiosaur in the continent and measures up to 1.75 meters (5.7 ft) in length. Megacephalosaurus was one of the largest marine reptiles of its time with an estimated length of 6–9 meters (20–30 ft). Its long snout and consistently sized teeth suggest that it preferred a diet of smaller-sized prey.
Luskhan is an extinct genus of brachauchenine pliosaur from the Cretaceous of Russia. The type and only species is Luskhan itilensis, named by Valentin Fischer and colleagues in 2017 from a well-preserved and nearly complete skeleton. As an early-diverging brachauchenine, Luskhan consequently exhibits an intermediate combination of traits seen in more basal and more derived pliosaurs. However, Luskhan departs significantly from other pliosaurs in that it exhibits a lack of adaptations in its skull to feeding on large prey; its slender snout, small teeth, and short tooth rows instead indicate a skull adapted for feeding on small, soft prey. With these features, it is the pliosaur that approaches closest to the distantly-related piscivorous polycotylids, having convergently evolved these traits more than 10 million years apart.
"Pliosaurus" andrewsi is an extinct species of pliosaurid plesiosaurs that lived during the Callovian stage of the Middle Jurassic, in what is now England. The only known fossils of this taxon were discovered in the Peterborough Member of the Oxford Clay Formation. Other attributed specimens have been discovered in various corners of Eurasia, but these are currently seen as indeterminate or coming from other taxa. The taxonomic history of this animal is quite complex, because several of its fossils were attributed to different genera of pliosaurids, before being concretely named and described in 1960 by Lambert Beverly Tarlo as a species of Pliosaurus. However, although the taxon was found to be valid, subsequent revisions found that it is not part of this genus, and therefore a taxonomic revision must be carried out on this species.
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