- Restoration of Styxosaurus and Xiphactinus
- Restoration
- Fossil skull and neck
Styxosaurus Temporal range: Late Cretaceous, | |
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Holotype skull of Styxosaurus snowii | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Superorder: | † Sauropterygia |
Order: | † Plesiosauria |
Family: | † Elasmosauridae |
Subfamily: | † Elasmosaurinae |
Genus: | † Styxosaurus Welles, 1943 |
Species | |
Synonyms | |
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Styxosaurus is a genus of plesiosaur of the family Elasmosauridae. Styxosaurus lived during the Campanian age of the Cretaceous period. Three species are known: S. snowii, S. browni, and S. rezaci.
The holotype specimen of Styxosaurus snowii a complete skull and 20 vertebrae and was first described by S.W. Williston [1] [2] Styxosaurus is named for the mythological River Styx (Στυξ), which separated the Greek underworld from the world of the living. The -saurus part comes from the Greek sauros (σαυρος), meaning "lizard" or "reptile." The type specimen was found on Hell Creek in Logan County, Kansas, which is the source of the genus name coined by Samuel Paul Welles, who described the genus in 1943. [3]
Another more complete specimen, SDSMT 451, was discovered near Iona, South Dakota, also in the US, in 1945. The specimen was originally described and named Alzadasaurus pembertoni by Welles and Bump (1949) and remained so until it was synonymized with S. snowii by Carpenter. [4] Its chest cavity contained about 250 gastroliths, or "stomach stones". Although it is mounted at the School of Mines as if its head were looking up and out of the water, such a position would be physically impossible. [5] In 2023, Elliott Smith and Robin O'Keefe also assigned this specimen to Styxosaurus but considered it to be indeterminate at the species level. [6]
In 2023, another species of Styxosaurus, S. rezaci, was named by Smith and O'Keefe, based on a specimen from the Cenomanian of Nebraska previously thought to belong to Thalassomedon . [6]
Styxosaurus was a large elasmosaur, with a long neck measuring about 5.25 metres (17.2 ft) in total. [4] [7] The S. snowii specimen NJSM 15435 was reported to measure 10 m (33 ft), though it lacks the skull and the frontmost neck vertebrae. [8] The cranium of the holotype measures 42 cm (1.38 ft) long, with its mandible measuring 48 cm (1.57 ft) long. [4] Its sharp teeth were conical and were adapted to puncture and hold rather than to cut; like other plesiosaurs, Styxosaurus swallowed its food whole. [9]
Styxosaurus snowii is from a group called elasmosaurs, and is closely related to Elasmosaurus platyurus , which was found in Kansas, USA, in 1867. [10]
The first Styxosaurus to be described was initially called Cimoliasaurus snowii by S.W. Williston in 1890. [1] The specimen included a complete skull and more than 20 cervical vertebrae ( KUVP 1301) that were found near Hell Creek in western Kansas by Judge E.P. West. [1]
The name was later changed to Elasmosaurus snowii by Williston in 1906 [11] and then to Styxosaurus snowii by Welles in 1943. [3]
A second species, Styxosaurus browni, was named by Welles in 1952. Although synonymized with Hydralmosaurus serpentinus by Kenneth Carpenter in 1999, it was revalidated by Rodrigo A. Otero in 2016. [4] [12]
The following cladogram shows the placement of Styxosaurus within Elasmosauridae following an analysis by Otero, 2016: [13]
Elasmosauridae |
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While most predators do not use gastroliths for grinding of food, almost all reasonably complete elasmosaur specimens include gastroliths. Although it is possible Styxosaurus may have used the stones as ballast, a Styxosaurus specimen found in the Pierre Shale of western Kansas included ground up fish bones mixed with the gastroliths. [14] In addition, the weight of the gastroliths found in elasmosaur specimens is always much less than 1% of the estimated weight of the living animal. [15]
While crocodiles and some other animals may use gastroliths for ballast today, it appears likely that elasmosaurs used them as a gastric mill. See Henderson (2006) contra Wings (2004). [16]
Styxosaurus, like most other plesiosaurs, probably fed on belemnites, fish ( Gillicus , etc.) and squid. With its interlocking teeth, Styxosaurus could grab on to its slippery prey before swallowing it.
Plesiosauroidea is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous periods. After their discovery, some plesiosauroids were said to have resembled "a snake threaded through the shell of a turtle", although they had no shell.
Elasmosaurus is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5 million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was sent to the American paleontologist Edward Drinker Cope, who named it E. platyurus in 1868. The generic name means "thin-plate reptile", and the specific name means "flat-tailed". Cope originally reconstructed the skeleton of Elasmosaurus with the skull at the end of the tail, an error which was made light of by the paleontologist Othniel Charles Marsh, and became part of their "Bone Wars" rivalry. Only one incomplete Elasmosaurus skeleton is definitely known, consisting of a fragmentary skull, the spine, and the pectoral and pelvic girdles, and a single species is recognized today; other species are now considered invalid or have been moved to other genera.
The Plesiosauria or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia.
Thalassomedon is a genus of plesiosaur, named by Welles in 1943.
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae.
Muraenosaurus is an extinct genus of cryptoclidid plesiosaur reptile from the Oxford Clay of Southern England. The genus was given its name due to the eel-like appearance of the long neck and small head. Muraenosaurus grew up to 5.2 metres (17 ft) in length and lived roughly between 160 Ma and 164 Ma in the Callovian of the middle Jurassic. Charles E. Leeds collected the first Muraenosaurus which was then described by H. G. Seeley. The specimen may have suffered some damage due to the casual style of Charles Leeds’ collection. The first muraenosaur was recovered with pieces missing from the skull and many of the caudal vertebrae absent. Because the animal was described from Charles Leeds’ collection it was given the name Muraenosaurus Leedsi. M. leedsi is the most complete specimen belonging to the genus Muraenosaurus and also the only species that is undoubtedly a member of the genus. Two other species have been tentatively referred to as members of the genus Muraenosaurus: M. reedii and Muraenosaurus beloclis Seeley 1892, which in 1910 became the separate genus Picrocleidus.
Libonectes is an extinct genus of sauropterygian reptile belonging to the plesiosaur order. It is known from specimens found in the Britton Formation of Texas (USA) and the Akrabou Formation of Morocco, which have been dated to the lower Turonian stage of the late Cretaceous period.
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species D. osborni and D. herschelensis, with two previous species having been assigned to new genera. Definitive specimens of D. osborni have been found in the late Coniacian to early Campanian rocks, while those of D. herschelensis have been found in the late Campanian to early Maastrichtian rocks. Dolichorhynchops was a prehistoric marine reptile measuring around 3 metres (9.8 ft) long. Its Greek generic name means "long-nosed face".
Brachauchenius is an extinct genus of pliosaurid that lived in North America and Morocco during the Late Cretaceous.
Tuarangisaurus is an extinct genus of elasmosaurid known from New Zealand. The type and only known species is Tuarangisaurus keyesi, named by "Pont" Wiffen, Joan Wiffen and Bill Moisley in 1986. The specific name honours Ian W. Keyes of the New Zealand Geological Survey.
Aphrosaurus was an extinct genus of plesiosaur from the Maastrichtian. The type species is Aphrosaurus furlongi, named by Welles in 1943. The holotype specimen was discovered in the Moreno Formation in Fresno County, California in 1939 by rancher Frank C. Piava. A second specimen - LACM 2832 - was also found in the same formation and initially diagnosed as a juvenile of the same species, but has since been removed from the genus.
The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico.
Aristonectes is an extinct genus of large elasmosaurid plesiosaurs that lived during the Maastrichtian stage of the Late Cretaceous. Two species are known, A. parvidens and A. quiriquinensis, whose fossil remains were discovered in what are now Patagonia and Antarctica. Throughout the 20th century, Aristonectes was a difficult animal for scientists to analyze due to poor fossil preparation, its relationships to other genera were uncertain. After subsequent revisions and discoveries carried out from the beginning of the 21st century, Aristonectes is now recognised as the type genus of the subfamily Aristonectinae, a lineage of elasmosaurids characterized by an enlarged skull and a reduced length of the neck.
Ogmodirus is an extinct genus of plesiosaur found in the Cenomanian-Turonian Greenhorn Limestone of Kansas. The type species, O. martini, was named by Samuel Wendell Williston and Roy Lee Moodie in 1913.
During the time of the deposition of the Niobrara Chalk, much life inhabited the seas of the Western Interior Seaway. By this time in the Late Cretaceous many new lifeforms appeared such as mosasaurs, which were to be some of the last of the aquatic lifeforms to evolve before the end of the Mesozoic. Life of the Niobrara Chalk is comparable to that of the Dakota Formation, although the Dakota Formation, which was deposited during the Cenomanian, predates the chalk by about 10 million years.
Albertonectes is an extinct genus of elasmosaurid plesiosaur known from the Late Cretaceous Bearpaw Formation of Alberta, Canada. It contains a single species, Albertonectes vanderveldei. Albertonectes is the longest elasmosaur, and more generally plesiosaur, known to date both in neck and total body length, estimated around 12 metres (39 ft) long and weighing up to 4.8 metric tons.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Nakonanectes bradti is an elasmosaurid plesiosaur of the late Cretaceous found in 2010 the state of Montana in the United States. It is one of the most recently known elasmosaurids to have lived in North America. Unlike other elasmosaurids, it has a relatively short neck.
Plesioelasmosaurus is an extinct genus of elasmosaurid plesiosaur from the Late Cretaceous Greenhorn Limestone of Kansas, United States. The genus contains a single species, P. walkeri, known from a partial skeleton.