A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In other species the rocks are ingested and pass through the digestive system and are frequently replaced. The grain size depends upon the size of the animal and the gastrolith's role in digestion. Other species use gastroliths as ballast. Particles ranging in size from sand to cobble have been documented.
Elasmosaurus is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5 million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was sent to the American paleontologist Edward Drinker Cope, who named it E. platyurus in 1868. The generic name means "thin-plate reptile", and the specific name means "flat-tailed". Cope originally reconstructed the skeleton of Elasmosaurus with the skull at the end of the tail, an error which was made light of by the paleontologist Othniel Charles Marsh, and became part of their "Bone Wars" rivalry. Only one incomplete Elasmosaurus skeleton is definitely known, consisting of a fragmentary skull, the spine, and the pectoral and pelvic girdles, and a single species is recognized today; other species are now considered invalid or have been moved to other genera.
Futabasaurus is a genus of plesiosaur from the Late Cretaceous of Fukushima, Japan. It was described and named in 2006, and was assigned to the family Elasmosauridae. The genus contains one species, F. suzukii.
Panoplosaurus is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first discovered in 1917, and named in 1919 by Lawrence Lambe, named for its extensive armour, meaning "well-armoured lizard". Panoplosaurus has at times been considered the proper name for material otherwise referred to as Edmontonia, complicating its phylogenetic and ecological interpretations, at one point being considered to have existed across Alberta, New Mexico and Texas, with specimens in institutions from Canada and the United States. The skull and skeleton of Panoplosaurus are similar to its relatives, but have a few significant differences, such as the lumpy form of the skull osteoderms, a completely fused shoulder blade, and regularly shaped plates on its neck and body lacking prominent spines. It was a quadrupedal animal, roughly 5 m (16 ft) long and 1,600 kg (3,500 lb) in weight. The skull has a short snout, with a very domed surface, and bony plates directly covering the cheek. The neck had circular groups of plates arranged around the top surface, both the forelimb and hindlimb were about the same length, and the hand may have only included three fingers. Almost the entire surface of the body was covered in plates, osteoderms and scutes of varying sizes, ranging from large elements along the skull and neck, to smaller, round bones underneath the chin and body, to small ossicles that filled in the spaces between other, larger osteoderms.
Libonectes is an extinct genus of sauropterygian reptile belonging to the plesiosaur order. It is known from specimens found in the Britton Formation of Texas (USA) and the Akrabou Formation of Morocco, which have been dated to the lower Turonian stage of the late Cretaceous period.
Umoonasaurus is an extinct genus of plesiosaur belonging to the family Leptocleididae. This genus lived approximately 115 million years ago during the Early Cretaceous period, in shallow seas covering parts of what is now Australia. It was a relatively small animal around 2.5 metres (8.2 ft) long. An identifying trait of Umoonasaurus is three crest-ridges on its skull.
Aphrosaurus was an extinct genus of plesiosaur from the Maastrichtian. The type species is Aphrosaurus furlongi, named by Welles in 1943. The holotype specimen was discovered in the Moreno Formation in Fresno County, California in 1939 by rancher Frank C. Piava. A second specimen - LACM 2832 - was also found in the same formation and initially diagnosed as a juvenile of the same species, but has since been removed from the genus.
Terminonatator is a genus of elasmosaurid plesiosaur from the Late Cretaceous of Saskatchewan, Canada. It is known from a skull and partial skeleton from a young adult, found in the Campanian-age Bearpaw Formation near Notukeu Creek in Ponteix. Terminonatator is currently one of the youngest plesiosaurs from the Western Interior Seaway.
Styxosaurus is a genus of plesiosaur of the family Elasmosauridae. Styxosaurus lived during the Campanian age of the Cretaceous period. Three species are known: S. snowii, S. browni, and S. rezaci.
Nichollssaura is an extinct genus of leptocleidid plesiosaur from the Early Cretaceous Boreal Sea of North America. The type species is N. borealis, found in the early Albian age Clearwater Formation near Fort McMurray, Alberta, Canada.
Aristonectes is an extinct genus of large elasmosaurid plesiosaurs that lived during the Maastrichtian stage of the Late Cretaceous. Two species are known, A. parvidens and A. quiriquinensis, whose fossil remains were discovered in what are now Patagonia and Antarctica. Throughout the 20th century, Aristonectes was a difficult animal for scientists to analyze due to poor fossil preparation, its relationships to other genera were uncertain. After subsequent revisions and discoveries carried out from the beginning of the 21st century, Aristonectes is now recognised as the type genus of the subfamily Aristonectinae, a lineage of elasmosaurids characterized by an enlarged skull and a reduced length of the neck.
Kaiwhekea is an extinct genus of plesiosaur from the Late Cretaceous of what is now New Zealand.
Brancasaurus is a genus of plesiosaur which lived in a freshwater lake in the Early Cretaceous of what is now North Rhine-Westphalia, Germany. With a long neck possessing vertebrae bearing distinctively-shaped "shark fin"-shaped neural spines, and a relatively small and pointed head, Brancasaurus is superficially similar to Elasmosaurus, albeit smaller in size at 3.26 metres (10.7 ft) in length as a subadult.
Tatenectes is a genus of cryptoclidid plesiosaur known from the Upper Jurassic of Wyoming. Its remains were recovered from the Redwater Shale Member of the Sundance Formation, and initially described as a new species of Cimoliosaurus by Wilbur Clinton Knight in 1900. It was reassigned to Tricleidus by Maurice G. Mehl in 1912 before being given its own genus by O'Keefe and Wahl in 2003. Tatenectes laramiensis is the type and only species of Tatenectes. While the original specimen was lost, subsequent discoveries have revealed that Tatenectes was a very unusual plesiosaur. Its torso had a flattened, boxy cross-section and its gastralia exhibit pachyostosis (thickening). The total length of Tatenectes has been estimated at 2–3 meters (6.6–9.8 ft).
Wapuskanectes is an extinct genus of early elasmosaurid known from Alberta, Canada, in the Albian aged Wabiskaw member of the Clearwater Formation.
Albertonectes is an extinct genus of elasmosaurid plesiosaur known from the Late Cretaceous Bearpaw Formation of Alberta, Canada. It contains a single species, Albertonectes vanderveldei. Albertonectes is the longest elasmosaur, and more generally plesiosaur, known to date both in neck and total body length, estimated around 12 metres (39 ft) long and weighing up to 4.8 metric tons.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Citipes is an extinct genus of caenagnathid theropod from the Late Cretaceous Dinosaur Park Formation in Alberta, Canada. The genus contains only one species, the type species, C. elegans. The generic name of Citipes is Latin for "fleet-footed", and the specific epithet "elegans" is Latin for "elegant". The type specimen of Citipes has a convoluted taxonomic history, and has been previously assigned to the genera Ornithomimus, Macrophalangia, Elmisaurus, Chirostenotes, and Leptorhynchos before being given its own genus in 2020.
Plesioelasmosaurus is an extinct genus of elasmosaurid plesiosaur from the Late Cretaceous Greenhorn Limestone of Kansas, United States. The genus contains a single species, P. walkeri, known from a partial skeleton.
Chubutinectes is an extinct genus of elasmosaurid plesiosaur from the Late Cretaceous La Colonia Formation of Argentina. The genus contains a single species, C. carmeloi, known from a partial skeleton and associated gastroliths.
