| Nothosauridae | |
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| Nothosaurus mirabilis fossil | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Superorder: | † Sauropterygia |
| Order: | † Nothosauroidea |
| Suborder: | † Nothosauria |
| Family: | † Nothosauridae Seeley, 1889 |
| Genera [1] | |
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Nothosauridae are an extinct family of carnivorous aquatic sauropterygian reptiles from the Triassic time period of China, France, Germany, Israel, Italy, Netherlands, Russia, Switzerland, and northern Africa. [2] [3]
The cladogram shown below follows Rieppel (1998)'s phylogenetic analysis of nothosaurids (bold terminal taxa contain living species). [1] Most of these relations are still considered correct today, [4] but despite Rieppel (1998)'s referral of Ceresiosaurus and Silvestrosaurus to Lariosaurus , some authors still consider them separate and many additional species have been named since this analysis. [5]
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A species level phylogenetic analysis of Nothosauridae was performed by Liu et al. (2014), and included all known valid species of the family apart from Lariosaurus stensioi (type of Micronothosaurus), Nothosaurus cymatosauroides , and Ceresiosaurus lanzi . The resultant topology is similar to the one obtained in Rieppel (1998) if the new additions are ignored, however this analysis found both Lariosaurus and Nothosaurus to be polyphyletic in regard to each-other and all other genera of the family, making a systematic revision of these two genera necessary. Below, their results are shown with type species of named nothosaurid genera noted. [6]
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Sauropterygia is an extinct taxon of diverse, aquatic reptiles that developed from terrestrial ancestors soon after the end-Permian extinction and flourished during the Triassic before all except for the Plesiosauria became extinct at the end of that period. The plesiosaurs would continue to diversify until the end of the Mesozoic. Sauropterygians are united by a radical adaptation of their pectoral girdle, adapted to support powerful flipper strokes. Some later sauropterygians, such as the pliosaurs, developed a similar mechanism in their pelvis. Uniquely among reptiles, sauropterygians moved their tail vertically like modern cetaceans and sirenians.
Nothosaurs were Triassic marine sauropterygian reptiles that may have lived like seals of today, catching food in water but coming ashore on rocks and beaches. They averaged about 3 metres (10 ft) in length, with a long body and tail. The feet were paddle-like, and are known to have been webbed in life, to help power the animal when swimming. The neck was quite long, and the head was elongated and flattened, and relatively small in relation to the body. The margins of the long jaws were equipped with numerous sharp outward-pointing teeth, indicating a diet of fish and squid.
Nothosaurus is an extinct genus of sauropterygian reptile from the Triassic period, approximately 240–210 million years ago, with fossils being distributed from North Africa and Europe to China. It is the best known member of the nothosaur order.
Pachypleurosaurs were primitive sauropterygian reptiles that vaguely resembled aquatic lizards, and were limited to the Triassic period. They were elongate animals, ranging in size from 20 cm to about a metre in length, with small heads, long necks, paddle-like limbs, and long deep tails. The limb girdles are greatly reduced, so it is unlikely these animals could move about on land. The widely spaced peg-like teeth project at the front of the jaws, indicate that these animals fed on fish.
Lariosaurus is an extinct genus of nothosaurid from the Middle Triassic of central and western Europe and China. With a length of just 60 centimetres (2.0 ft), it was one of the smallest known nothosaurs. First discovered at Perledo on the Lake Como in 1830, it was named in 1847 by Curioni, its name meaning "Lizard from Larius", the ancient name of the lake. This makes it one of the earliest studied reptiles from the Alps. It is known from an almost complete skeleton holotype and several other fairly complete fossils.
Ceresiosaurus is an extinct aquatic genus of lariosaurine nothosaurid sauropterygian known from the Middle Triassic of Monte San Giorgio, southern Switzerland and northern Italy. Ceresiosaurus, meaning "Lizard of Ceresio". The type species, Ceresiosaurus calcagnii, was named by Bernhard Peyer in 1931. C. calcagnii is known from both the Cava superiore and Cava inferiore beds of the Lower Meride Limestone at Monte San Giorgio, dating to the latest Anisian of the Middle Triassic. Rieppel (1998) suggested that the back then monospecific genus Ceresiosaurus, is a junior synonym of the better known Lariosaurus, yet he kept it type species as a separate species under the new combination L. calcagnii. In 2004, however, this synonymy was objected by Hänni who described and name a second species of Ceresiosaurus, C. lanzi - a separation supported by several other authors since. This species is known only from the stratigraphically younger Cassima beds of Monte San Giorgio, although also from the Lower Meride Limestone, dating to possibly the lowest Ladinian age. The species in a subtropical lagoonal environment with varying open marine influences, and alongside many related but smaller species of nothosaurids and pachypleurosaurids. Ceresiosaurus represents one of the largest vertebrate of up to 3 m snout-tail length from the very diversified paleoenvironment of the Middle Triassic Monte San Giorgio.
Pistosaurus is an extinct genus of aquatic sauropterygian reptile closely related to plesiosaurs. Fossils have been found in France and Germany, and date to the Middle Triassic. It contains a single species, Pistosaurus longaevus. Pistosaurus is known as the oldest "subaquatic flying" reptile on earth.
Cyamodus is a genus of placodonts discovered from fossil remains in Germany in the early-to-mid-19th century and was named by Christian Erich Hermann von Meyer in 1863. The fossils have been dated to the Triassic Period, from the Anisian to Ladinian stages. Considered a possible turtle ancestor due to its wide flattened torso covered in dermal ossicles, but now is considered to only be distantly related. Derived from a sister to Paraplacodus, Cyamodus phylogenetically preceded Placochelys.
Thalattosauria is an extinct order of prehistoric marine reptiles that lived in the middle to late Triassic period. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.
Helveticosaurus is an extinct genus of diapsid marine reptile known from the Middle Triassic of southern Switzerland. It contains a single species, Helveticosaurus zollingeri, known from the nearly complete holotype T 4352 collected at Cava Tre Fontane of Monte San Giorgio, an area well known for its rich record of marine life during the Middle Triassic.
Sinosaurosphargis is an extinct genus of basal marine saurosphargid reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a single species, Sinosaurosphargis yunguiensis.
Wumengosaurus is an extinct aquatic reptile from the Middle Triassic Guanling Formation of Guizhou, southwestern China. It was originally described as a basal eosauropterygian and usually is recovered as such by phylogenetic analyses, although one phylogeny has placed it as the sister taxon to Ichthyosauromorpha while refraining from a formal re-positioning.
Largocephalosaurus is an extinct genus of basal saurosphargid, a marine reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a type species, Largocephalosaurus polycarpon, and a second species L. qianensis.
Hanosaurus is an extinct genus of marine reptiles that existed during the Triassic period in what is now China. It was described by Young in 1972, and the type species is Hanosaurus hupehensis. Its affinities are unclear; it has been both described as a member of the Pachypleurosauria and as a sister group of that group. It was a small animal that likely fed on soft-bodied prey.
Wangosaurus is an extinct genus of basal pistosauroid known from the Middle Triassic Falang Formation of Xingyi in Guizhou Province, southwestern China. It contains a single species, Wangosaurus brevirostris, first described and named by Le-Tian Ma, Da-Yong Jiang, Olivier Rieppel, Ryosuke Motani and Andrea Tintori in 2015. The specific name brevirostris comes from Greek for "short snout". It is known solely from its holotype, a nearly complete and articulated skeleton missing only the rear part of its tail.
Germanosaurus, meaning "German lizard", is an extinct aquatic genus of nothosaurid sauropterygian known from the early Middle Triassic Lower Muschelkalk of what was known as Upper Silesia, now a part of Poland. The type species of Germanosaurus is G. latissimus, originally named as a species of Nothosaurus. After a new generic name was erected for it, the holotype fragmentary skull was lost, possibly during World War II. Rieppel (1997) thus considered the species to be a nomen dubium in the species. However, from surviving illustrations and descriptions of the material, he concluded that another taxon known as Cymatosaurus schafferi, is referable to Germanosaurus and possibly even represents the same species as G. latissimus. Rieppel removed the species from Cymatosaurus and created the new combination G. schafferi, making it the only valid species of Germanosaurus diagnosable to the species level.
Silvestrosaurus is an extinct aquatic genus of lariosaurine nothosaurid sauropterygian known from the Middle Triassic of Monte San Giorgio, southern Switzerland. It contains a single species, Silvestrosaurus buzzii, originally considered to be a species of the closely related Lariosaurus. The species was named by Tschanz in 1989, based solely on the holotype PIMUZ T/2804 comprising the skull, the lower jaw, and a dis-articulated partial postcranial skeleton. Cyamodus hildegardis tooth bearing elements were found in the stomach region of the specimen. The holotype was collected at Punkt 902 of Monte San Giorgio, from layer 97 of the Grenzbitumen zone, dating to the Anisian-Ladinian boundary of the Middle Triassic. Kuhn-Schnyner (1990) reassigned the species to its own genus, creating the combination S. buzzii. The generic name honors a church near the collection locality of the holotype, dedicated to Saint Sylvester, a Pope during the reign of Constantine the Great, and from Greek saurus, meaning "lizard", a common suffix for genus names of extinct reptile.
Saurosphargidae is an extinct family of marine reptiles known from the early Middle Triassic of Europe and China.
Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi.
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