Eoplesiosaurus is known only from the holotype specimen TTNCM8348, a complete and articulated postcranialskeleton. It preserved three-dimensionally with some gastralia presented. It was collected at Watchet, of Somerset, from the Pre-Planorbis beds of the Blue Lias Formation of the Lower Lias Group. These beds likely occur below the first occurrence of the ammonite Psiloceras planorbis. Thus, they probably fall within the earliest Hettangian P. tilmanni Chronozone, which is about 199.6-198million years old, immediately following the Triassic–Jurassic Boundary. Plesiosaurs fossils which were discovered at Street (e.g. Avalonnectes and Thalassiodracon), or at penecontemporaneous strata such as the Watchet locality, represent the earliest known occurrence of the Plesiosauria. Hence, Eoplesiosaurus is the oldest plesiosauroid and one of the oldest plesiosaurs, to date.[1]
Eoplesiosaurus is a proportionally long-necked plesiosaurian with at least 38 cervical vertebrae. The presence of small, conical lateral projections on the bases of the anterior cervical prezygapophyses represents its autapomorphy.
Classification
A phylogenetic analysis performed by Benson et al. (2012) found it to be the basalmost known plesiosauroid. The cladogram below shows Eoplesiosaurus phylogenetic position among other plesiosaurs following Benson et al. (2012).[1]
Plesiosauroidea is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous periods. After their discovery, some plesiosauroids were said to have resembled "a snake threaded through the shell of a turtle", although they had no shell.
Plesiosaurus is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias of England. It is distinguishable by its small head, long and slender neck, broad turtle-like body, a short tail, and two pairs of large, elongated paddles. It lends its name to the order Plesiosauria, of which it is an early, but fairly typical member. It contains only one species, the type, Plesiosaurus dolichodeirus. Other species once assigned to this genus, including P. brachypterygius, P. guilielmiimperatoris, and P. tournemirensis have been reassigned to new genera, such as Hydrorion, Seeleyosaurus and Occitanosaurus.
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled pleisiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used.
Macroplata is an extinct genus of Early Jurassic rhomaleosaurid plesiosaur which grew up to 4.65 metres (15.3 ft) in length. Like other plesiosaurs, Macroplata probably lived on a diet of fish, using its sharp needle-like teeth to catch prey. Its shoulder bones were fairly large, indicating a powerful forward stroke for fast swimming. Macroplata also had a relatively long neck, twice the length of the skull, in contrast to pliosaurs.
Thalassiodracon (tha-LAS-ee-o-DRAY-kon) is an extinct genus of plesiosauroid from the Pliosauridae that was alive during the Late Triassic-Early Jurassic (Rhaetian-Hettangian) and is known exclusively from the Lower Lias of England. The type and only species, is Thalassiodracon (Plesiosaurus) hawkinsi.
Colymbosaurus is a genus of cryptoclidid plesiosaur from the Late Jurassic (Callovian-Tithonian) of the UK and Svalbard, Norway. There are two currently recognized species, C. megadeirus and C. svalbardensis. Both species are relatively medium-sized plesiosaurs, reaching 5 metres (16 ft) in length and 700 kilograms (1,500 lb) in body mass.
Archaeonectrus is an extinct genus of pliosaur from the Early Jurassic (Sinemurian) of what is now southeastern England. The type species is Archaeonectrusrostratus, first named by Sir Richard Owen in 1865, which was moved to its own genus by N.I. Novozhilov in 1964. It was a relatively small plesiosaur, measuring 3.67 m (12.0 ft) long.
The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassic times, between 195 and 200 million years ago. The Blue Lias is famous for its fossils, especially ammonites.
Eurycleidus is an extinct genus of large-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, E. arcuatus. Like other plesiosaurs, Eurycleidus probably lived on a diet of fish, using its sharp needle-like teeth to catch prey. Its shoulder bones were fairly large, indicating a powerful forward stroke for fast swimming.
Plesiopterys is an extinct genus of plesiosaur originating from the Posidonienschiefer of Holzmaden, Germany, and lived during the Early Jurassic period. It is thought to be the sister taxon to all other plesiosauroids including the Plesiosaurus, and is placed outside of the Plesiosauroidea group. Plesiopterys wildi is the one known species within the genus, and is 220 centimeters long, or about 7.2 feet, and its body and skull are both relatively small. It possesses a unique combination of both primitive and derived characters, and is currently displayed at the State Museum of Natural History, Germany.
Rhomaleosauridae is a family of plesiosaurs from the Earliest Jurassic to the latest Middle Jurassic of Europe, North America, South America and possibly Asia. Most rhomaleosaurids are known from England, many specifically from lower Blue Lias deposits that date back to the earliest Jurassic, just at the boundary with the Triassic. In fact, to date only two undisputed rhomaleosaurids were named from outside Europe - the closely related Borealonectes russelli and Maresaurus coccai from Canada and Argentina, respectively. These two species are also the only Middle Jurassic representatives of the family. Rhomaleosauridae was formally named by Kuhn in 1961, originally proposed to include Rhomaleosaurus cramptoni and its relatives, which have short necks and large heads relatively to plesiosauroids like Elasmosaurus and Plesiosaurus, but longer necks and smaller heads relatively to advanced pliosaurids like Pliosaurus and Kronosaurus.
Meyerasaurus is an extinct genus of rhomaleosaurid known from Holzmaden, Baden-Württemberg of southwestern Germany.
Avalonnectes is an extinct genus of small-bodied rhomaleosaurid known from the Early Jurassic period of the United Kingdom. It contains a single species, A. arturi.
Stratesaurus is an extinct genus of small-bodied rhomaleosaurid plesiosaur known from the Early Jurassic period of the United Kingdom. It contains a single species, S. taylori. It was a small plesiosaur, with a skull length of 18 cm (7.1 in) and a body length of 2 m (6.6 ft).
Microcleididae is an extinct family of basal plesiosauroid plesiosaurs from the Early Jurassic of France, Germany, Portugal and the United Kingdom. Currently, the oldest and the most known microcleidid is Eretmosaurus from the middle Sinemurian of the United Kingdom. Microcleididae was formally named and described by Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller in 2012.
Anningasaura is an extinct genus of basal plesiosaur. It is known from a single type species, A. lymense, discovered in Early Jurassic rocks of Lyme Regis in the United Kingdom.
Spitrasaurus is an extinct genus of cryptoclidid plesiosauroid plesiosaur known from the uppermost Jurassic of central Spitsbergen, Norway and likely also Kimmeridge, England. It is named after a syllabic abbreviation for Spitsbergen Travel.
This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptiles that flourished during the Mesozoic Era. The first scientifically documented plesiosaur fossils were discovered during the early 19th century by Mary Anning. Plesiosaurs were actually discovered and described before dinosaurs. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial reptiles. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors.
Atychodracon is an extinct genus of rhomaleosaurid plesiosaurian known from the Late Triassic - Early Jurassic boundary of England. It contains a single species, Atychodracon megacephalus, named in 1846 originally as a species of Plesiosaurus. The holotype of "P." megacephalus was destroyed during a World War II air raid in 1940 and was later replaced with a neotype. The species had a very unstable taxonomic history, being referred to four different genera by various authors until a new genus name was created for it in 2015. Apart from the destroyed holotype and its three partial casts, a neotype and two additional individuals are currently referred to Atychodracon megacephalus, making it a relatively well represented rhomaleosaurid.
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