Trilophosaurus (Greek for "lizard with three ridges") is a lizard-like trilophosauridallokotosaur known from the Late Triassic of North America. It was a herbivore up to 2.5 m long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak.
T. buettneri.T. buettneri compared to a human.
The skull is also unusual in that the lower temporal opening is missing, giving the appearance of a euryapsid skull.[citation needed] Because of this, the trilophosaurs were once classified with placodonts within Sauropterygia. Carroll (1988) suggested that the lower opening may have been lost to strengthen the skull.
Life reconstruction of Trilophosaurus buettneri
Trilophosaurus is traditionally thought to include two valid species: the typical T. buettneri and the more robust T. jacobsi. In 1993, paleontologists Hans-Dieter Sues and Paul E. Olsen reassigned T. jacobsi, as well as two additional trilophosaurids (Tricuspisaurus and Variodens), to Procolophonidae based on similarities between its tricuspid teeth and those of the newly described procolophonid Xenodiphyodon.[1] This view persisted in subsequent publications[2] until the cranial material of T. jacobsi was described by Heckert et al. (2006). The new material confirmed the originally classification that T. jacobsi, as well as Tricuspisaurus and Variodens based on similarities to it, are indeed trilophosaurids.[3] Meanwhile, a third species of Trilophosaurus, T. dornorum, was named by Mueller & Parker (2006) based on teeth of a robust individual.[4] However, Spielmann et al. (2009) argued that the robustness of the new species is not sufficient to differentiate it from other Trilophosaurus species, especially in light of new robust specimens of T. jacobsi. Therefore, they considered T. dornorum to be a junior synonym of T. jacobsi,[5] a view that was maintained since in other publications.[6] Spielmann et al. (2006) redescribed the type material of Malerisaurus langstoni and concluded that it's indistinguishable from T. buettneri, and thus M. langstoni represents its synonym.[7]
Nesbitt et al. (2015) performed a phylogenetic analysis focusing on relations within Allokotosauria and recovered T. jacobsi to be more closely related to Spinosuchus caseanus than to the type species of Trilophosaurus. To further test this possibility, the types of S. caseanus and T. jacobsi were scored separately from the referred the Kahle Trilophosaurus Quarry elements (referred to T. jacobsi by Spielmann et al. (2008) or to S. caseanus by Spielmann et al. (2009)). A phylogenetic analysis recovered the three in a monophyletic clade to the exclusion of T. buettneri based on a single autapomorphy. Furthermore, the types of S. caseanus and T. jacobsi as well as the Kahle Quarry material all scored identically, suggesting that T. jacobsi not only should be reassigned to Spinosuchus, but in fact represents the junior synonym of its type and only species (S. caseanus). Nesbitt et al. (2015) refrained from officially synonymizing the two taxa pending further study of other advanced trilophosaurids.[6]
Sources
Benton, M. J. (2000), Vertebrate Paleontology, 2nd ed. Blackwell Science Ltd, p. 144
Carroll, R. L. (1988), Vertebrate Paleontology and Evolution, W.H. Freeman & Co. p. 266
↑ Sues, H.-D.; Olsen, P.E. (1993). "A new procolophonid and a new tetrapod of uncertain, possibly procolophonian affinities from the Upper Triassic of Virginia". Journal of Vertebrate Paleontology. 13 (3): 282–286. doi:10.1080/02724634.1993.10011510.
1 2 Nesbitt, S.J.; Flynn, J.J.; Pritchard, A.C.; Parrish, M.J.; Ranivoharimanana, L.; Wyss, A.R. (2015). "Postcranial osteology of Azendohsaurus madagaskarensis (?Middle to Upper Triassic, Isalo Group, Madagascar) and its systematic position among stem archosaur reptiles". Bulletin of the American Museum of Natural History (398): 1–126. doi:10.5531/sd.sp.15. hdl:2246/6624. ISSN0003-0090.
Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.
Azendohsaurus is an extinct genus of herbivorous archosauromorph reptile from roughly the late Middle to early Late Triassic Period of Morocco and Madagascar. The type species, Azendohsaurus laaroussii, was described and named by Jean-Michel Dutuit in 1972 based on partial jaw fragments and some teeth from Morocco. A second species from Madagascar, A. madagaskarensis, was first described in 2010 by John J. Flynn and colleagues from a multitude of specimens representing almost the entire skeleton. The generic name "Azendoh lizard" is for the village of Azendoh, a local village near where it was first discovered in the Atlas Mountains. It was a bulky quadruped that unlike other early archosauromorphs had a relatively short tail and robust limbs that were held in an odd mix of sprawled hind limbs and raised forelimbs. It had a long neck and a proportionately small head with remarkably sauropod-like jaws and teeth.
Revueltosaurus is an extinct genus of suchian pseudosuchian from Late Triassic deposits of New Mexico, Arizona and North Carolina, United States. Many specimens, mostly teeth, have been assigned to Revueltosaurus over the years. Currently, three species are included in this genus, all of which were originally thought to represent monospecific genera of basal ornithischian dinosaurs. Revueltosaurus was about 1 meter long.
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs first appeared in the Early Triassic, reaching their broadest abundance and a global distribution during the Carnian stage of the Late Triassic.
Trilophosaurs are lizard-like Triassic allokotosaur reptiles related to the archosaurs. The best known genus is Trilophosaurus, a herbivore up to 2.5 metres long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak.
Spinosuchus is an extinct genus of trilophosaurid allokotosaur from the Late Triassic of Texas, southern United States. It has been assigned to a variety of groups over its history, from coelophysid dinosaur to pseudosuchian to uncertain theropod dinosaur and to Proterosuchidae. This uncertainty is not unusual, given that it was only known from a poorly preserved, wall-mounted, partial vertebral column of an animal that lived in a time of diverse, poorly known reptile groups. However, newly collected material and recent phylogenetic studies of early archosauromorphs suggest that it represents an advanced trilophosaurid very closely related to Trilophosaurus.
Malerisaurus is an extinct genus of archosauromorph known from Andhra Pradesh of India and Texas of the USA.
The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico. It is one of several formations encompassed by the Dockum Group.
The Chinle Formation is an extensive geological unit in the southwestern United States, preserving a very diverse fauna of Late Triassic animals and plants. This is a list of fossilized organisms recovered from the formation.
Uatchitodon is an extinct genus of Late Triassic reptile known only from isolated teeth. Based on the structure of the teeth, Uatchitodon was probably a carnivorous archosauromorph. Folded grooves on the teeth indicate that the animal was likely venomous, with the grooves being channels for salivary venom. The teeth are similar to those of living venomous squamates such as Heloderma and venomous snakes. Uatchitodon is the earliest known venomous reptile.
Variodens is an extinct genus of trilophosaur. Fossils have been found from the Emborough Quarries in the Mendip Hills of Somerset, England. These fossils have been uncovered from a Late Triassic fissure fill within Carboniferous-age limestone. The type and only known species is V. inopinatus, named in 1957.
Tricuspisaurus is an extinct genus of reptile originally described as a trilophosaurid; it was later considered likely to be a procolophonid, but recent analyses have affirmed the original classification. Fossils are known from the Ruthin Quarry in Glamorgan, Wales, one of several Late Triassic to Early Jurassic British fissure deposits. Like some trilophosaurs, it has an edentulous, or toothless beak. Tricuspisaurus gets its name from its heterodont dentition, which includes tricuspid teeth, or teeth with three cusps. The type species, T. thomasi, was named in 1957 along with the possible trilophosaur Variodens inopinatus from Somerset, England.
Coelodontognathus is an extinct genus of reptile from the Early Triassic of European Russia. It was originally described as a procolophonid parareptile in 1967 but was reclassified as a possible trilophosaurid archosauromorph in 2008. The genus includes two species: the type species C. donensis and C. ricovi. C. donensis is known from the holotype PIN 4173/129 and the referred PIN 4173/130, and C. ricovi is known from the holotype PIN 4173/127 and the referred PIN 4173/128, all of which represent dentaries that are housed at the Paleontological Institute, Russian Academy of Sciences. Another dentary, SGU 104/3105, originally referred to C. donensis was reassigned to its own genus and species Vitalia grata by Ivakhnenko, 1973. The fossils have been found at the Donskaya Luka Locality near the village of Sirotinskaya and the Don River in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. Like Coelodontognathus, Vitalia which is known from the same locality was also first identified as a procolophonid and later reclassified as a trilophosaurid. Coelodontognathus and Vitalia are similar to procolophonids in that they have wide teeth but differs from them in that they have tooth roots set deep into the jaws.
Pamelaria is an extinct genus of allokotosaurian archosauromorph reptile known from a single species, Pamelaria dolichotrachela, from the Middle Triassic of India. Pamelaria has sprawling legs, a long neck, and a pointed skull with nostrils positioned at the very tip of the snout. Among early archosauromorphs, Pamelaria is most similar to Prolacerta from the Early Triassic of South Africa and Antarctica. Both have been placed in the family Prolacertidae. Pamelaria, Prolacerta, and various other Permo-Triassic reptiles such as Protorosaurus and Tanystropheus have often been placed in a group of archosauromorphs called Protorosauria, which was regarded as one of the most basal group of archosauromorphs. However, more recent phylogenetic analyses indicate that Pamelaria and Prolacerta are more closely related to Archosauriformes than are Protorosaurus, Tanystropheus, and other protorosaurs, making Protorosauria a polyphyletic grouping.
The Colorado City Formation is a Late Triassic geologic formation in the Dockum Group of Texas, United States. It has previously been known as the Iatan Member, Colorado City Member or 'Pre-Tecovas Horizon'.
Allokotosauria is a clade of early archosauromorph reptiles from the Middle to Late Triassic known from Asia, Africa, North America and Europe. Allokotosauria was first described and named when a new monophyletic grouping of specialized herbivorous archosauromorphs was recovered by Sterling J. Nesbitt, John J. Flynn, Adam C. Pritchard, J. Michael Parrish, Lovasoa Ranivoharimanana and André R. Wyss in 2015. The name Allokotosauria is derived from Greek meaning "strange reptiles" in reference to unexpected grouping of early archosauromorph with a high disparity of features typically associated with herbivory.
Azendohsauridae is a family of allokotosaurian archosauromorphs that lived during the Middle to Late Triassic period, around 242-216 million years ago. The family was originally named solely for the eponymous Azendohsaurus, marking out its distinctiveness from other allokotosaurs, but as of 2022 the family now includes four other genera: the basal genus Pamelaria, the large horned herbivore Shringasaurus, and two carnivorous genera grouped into the subfamily-level subclade Malerisaurinae, Malerisaurus and Puercosuchus, and potentially also the dubious genus Otischalkia. Most fossils of azendohsaurids have a Gondwanan distribution, with multiple species known across Morocco and Madagascar in Africa as well as India, although fossils of malerisaurine azendohsaurids have also been found in the southwestern United States of North America.
Doniceps is an extinct genus of reptile from the Early Triassic of European Russia known from the type species D. lipovensis. It was solely known from the holotype premaxilla 104/3106 housed at Saratov University, however the specimen is probably lost. It was collected at the Donskaya Luka Locality near the village of Sirotinskaya in Ilovlinsky District, Volgograd Oblast, from the Lipovskaya Formation of the Gamskii Horizon. The generic name refers to the nearby Don River added the Greek suffix for "head", -ceps. The specific name is derived from the name of the type locality Donskaya Luka, also known as Lipovaya Balka. Named by Otschev and Rykov in 1968 as an archosauromorph, Arkhangelskii & Sennikov (2008) classified the taxon as a possible trilophosaurid. Doniceps is thought to be similar to Coelodontognathus and Vitalia, both of which are known exclusively from the same locality and were first identified as procolophonids but later reclassified as trilophosaurids.
Oryctorhynchus is an extinct genus of rhynchosaur from the Late Triassic (Carnian-Norian)-aged Wolfville Formation of Nova Scotia, Canada that may have been the same animal as Beesiiwo. The type species, O. bairdi, was named and described in 2020. It was originally seen as a species of Hyperodapedon until 2020.
Puercosuchus is an extinct genus of archosauromorph reptile from the Late Triassic (Norian) of what is now Arizona, North America. It includes only the type species P. traverorum, and was described and named in 2022. Puercosuchus is known mainly from two bonebeds in the Blue Mesa Member of the Chinle Formation, preserving the mixed remains of multiple individuals in each one representing almost the entire skeleton. It is a member of the Azendohsauridae, a clade of Triassic reptiles that was initially recognised by adaptations for herbivory. However, Puercosuchus and its close relatives in the subclade Malerisaurinae retained the carnivorous diet and body form ancestral to archosauromorphs. Unlike non-malerisaurine azendohsaurids, Puercosuchus had a long and shallow snout with sharp, blade-like teeth similar to those of carnivorous dinosaurs. Despite its seemingly ancestral morphology and ecology, Puercosuchus is the youngest known genus of azendohsaurid in the world. The discovery of Puercosuchus allowed palaeontologists to recognise similar bones and teeth that had been collected from Late Triassic southwestern North America in the past as belonging to it or similar animals, acting as a sort of "rosetta stone" for malerisaurine azendohsaurid anatomy.
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