Spinosuchus Temporal range: Late Triassic, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Clade: | † Allokotosauria |
Order: | † Trilophosauria |
Family: | † Trilophosauridae |
Genus: | † Spinosuchus von Huene, 1932 |
Species | |
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Synonyms | |
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Spinosuchus (meaning "spined crocodile") is an extinct genus of trilophosaurid allokotosaur from the Late Triassic of Texas, southern United States. It has been assigned to a variety of groups over its history, from coelophysid dinosaur to pseudosuchian to uncertain theropod dinosaur and to Proterosuchidae. This uncertainty is not unusual, given that it was only known from a poorly preserved, wall-mounted, partial vertebral column of an animal that lived in a time of diverse, poorly known reptile groups. However, newly collected material and recent phylogenetic studies of early archosauromorphs suggest that it represents an advanced trilophosaurid very closely related to Trilophosaurus . [1] [2]
In 1922, Ermine Cowles Case described a partial vertebral column (UMMP 7507) he'd discovered in 1921 from the Tecovas Member of the Carnian-age Upper Triassic Dockum Formation of Crosby County, Texas, as Coelophysis sp. (Coelophysis at that time also being poorly known). [3] He considered it to be about 2.5 meters (8.5 ft) long. [3] Additional material was referred to it, including a femur (UMMP 3396), [3] an ilium (UMMP 8870), and a basicranium (UMMP 7473). [4] These additional remains have since been recognized as belonging to a variety of other Triassic animals, all of which were poorly known or unknown at the time: the femur to an aetosaur, possibly Desmatosuchus , [5] the ilium to a herrerasaurid, either Chindesaurus or Caseosaurus , depending on the taxonomic authority, [6] and the basicranium to the rauisuchian Postosuchus . [7]
Friedrich von Huene recognized it as a new genus in 1932, and named it in honor of Case. [8] He considered it to be a "podokesaurid". [8] Since the 1970s, though, it has been considered a nondinosaurian. [9] [10] [11] However, a review by Hunt et al. in 1998 suggested that it was a theropod, possibly a herrerasaurid, citing its hollow centra as evidence for dinosaurian affinities. [5] In an abstract for the 1999 Society of Vertebrate Paleontology meeting and his unpublished thesis, Richards recognized that it had a variety of characters that are apomorphic for various dinosaur subgroups, but that these are also found in different basal archosaur groups, and that the poorly preserved, distorted, and reconstructed vertebrae offer no evidence for assignment to any major archosaur group; it does, though, show some resemblances to the trilophosaurs. [12] [13] Further review, as part of a larger series of papers on the evolution of dinosaurs in the Late Triassic by Sterling Nesbitt, Randall Irmis, and William Parker, found Spinosuchus to be a valid genus. However, the authors could not classify it beyond Archosauriformes, and disagreed with Richards' trilophosaur hypothesis. [14] Justin Spielmann and colleagues published a redescription in 2009 that concluded Spinosuchus was a trilophosaurid closely related to Trilophosaurus . [1]
Nesbitt et al. (2015) performed a phylogenetic analysis focusing on relations within Allokotosauria and recovered Trilophosaurus jacobsi to be more closely related to S. caseanus than to the type species of Trilophosaurus, Trilophosaurus buettneri . To further test this possibility, the types of S. caseanus and T. jacobsi were scored separately from the referred Kahle Trilophosaurus Quarry elements (referred to T. jacobsi by Spielmann et al. (2008) or to S. caseanus by Spielmann et al. (2009)). A phylogenetic analysis recovered the three in a monophyletic clade to the exclusion of T. buettneri based on a single autapomorphy. Furthermore, the types of S. caseanus and T. jacobsi as well as the Kahle Quarry material all scored identically, suggesting that T. jacobsi not only should be reassigned to Spinosuchus, but in fact represents the junior synonym of its type and only species (S. caseanus). Nesbitt et al. (2015) refrained from officially synonymizing the two taxa pending further study of other advanced trilophosaurids. [2]
Coelophysis is a genus of coelophysid theropod dinosaur that lived approximately 215 to 208.5 million years ago during the Late Triassic period from the middle to late Norian age in what is now the southwestern United States. Megapnosaurus was once considered to be a species within this genus, but this interpretation has been challenged since 2017 and the genus Megapnosaurus is now considered valid.
Archosauromorpha is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs rather than lepidosaurs. Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period.
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.
Postosuchus, meaning "Crocodile from Post", is an extinct genus of rauisuchid reptiles comprising two species, P. kirkpatricki and P. alisonae, that lived in what is now North America during the Late Triassic. Postosuchus is a member of the clade Pseudosuchia, the lineage of archosaurs that includes modern crocodilians. Its name refers to Post Quarry, a place in Texas where many fossils of the type species, P. kirkpatricki, were found. It was one of the apex predators of its area during the Triassic, larger than the small dinosaur predators of its time. It was a hunter which probably preyed on large bulky herbivores like dicynodonts and many other creatures smaller than itself.
Gojirasaurus is a potentially dubious genus of coelophysoid theropod dinosaur named after the giant monster movie character Godzilla.
Shuvosaurus is a genus of beaked, bipedal poposauroid pseudosuchian from the Late Triassic of western Texas. Despite superficially resembling a theropod dinosaur, especially the ostrich-like ornithomimids, it is instead more closely related to living crocodilians than to dinosaurs. Shuvosaurus is known by the type and only species S. inexpectatus, and is closely related to the very similar Effigia within the clade Shuvosauridae. Shuvosaurus was originally described from a restored skull and very few fragmentary postcranial bones as a probable ornithomimosaur, or at least a very ornithomimosaur-like early theropod. The true pseudosuchian affinities of Shuvosaurus were only recognised after the discovery of Effigia linked the skull of Shuvosaurus with similar poposauroid skeletal remains found in the same quarry.
Chindesaurus is an extinct genus of basal saurischian dinosaur from the Late Triassic of the southwestern United States. It is known from a single species, C. bryansmalli, based on a partial skeleton recovered from Petrified Forest National Park in Arizona. The original specimen was nicknamed "Gertie", and generated much publicity for the park upon its discovery in 1984 and airlift out of the park in 1985. Other fragmentary referred specimens have been found in Late Triassic sediments throughout Arizona, New Mexico, and Texas, but these may not belong to the genus. Chindesaurus was a bipedal carnivore, approximately as large as a wolf.
Trilophosaurs are lizard-like Triassic allokotosaur reptiles related to the archosaurs. The best known genus is Trilophosaurus, a herbivore up to 2.5 metres long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak.
Poposaurus is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, and Texas. Except for the skull, most parts of the skeleton are known. The type species, P. gracilis, was described and named by Maurice Goldsmith Mehl in 1915. A second species, P. langstoni, was originally the type species of the genus Lythrosuchus. Since it was first described, Poposaurus has been variously classified as a dinosaur, a phytosaur, and a "rauisuchian".
Dinosauromorpha is a clade of avemetatarsalians that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, with later formulations specifically excluding pterosaurs from the group. Birds are the only dinosauromorphs which survive to the present day.
Erythrosuchus is an extinct genus of archosauriform reptiles from the Triassic of South Africa. Remains have been found from the Cynognathus Assemblage Zone of the Beaufort Group in the Karoo of South Africa.
Trilophosaurus is a lizard-like trilophosaurid allokotosaur known from the Late Triassic of North America. It was a herbivore up to 2.5 m long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak.
The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico. It is one of several formations encompassed by the Dockum Group.
Tawa is a genus of possible basal theropod dinosaurs from the Late Triassic period. The fossil remains of Tawa hallae, the type and only species were found in the Hayden Quarry of Ghost Ranch, New Mexico, US. Its discovery alongside the relatives of Coelophysis and Herrerasaurus supports the hypothesis that the earliest dinosaurs arose in Gondwana during the early Late Triassic period in what is now South America, and radiated from there around the globe. The specific name honours Ruth Hall, founder of the Ghost Ranch Museum of Paleontology.
Variodens is an extinct genus of trilophosaur. Fossils have been found from the Emborough Quarries in the Mendip Hills of Somerset, England. These fossils have been uncovered from a Late Triassic fissure fill within Carboniferous-age limestone. The type and only known species is V. inopinatus, named in 1957.
Tricuspisaurus is an extinct genus of reptile originally described as a trilophosaurid; it was later considered likely to be a procolophonid, but recent analyses have affirmed the original classification. Fossils are known from the Ruthin Quarry in Glamorgan, Wales, one of several Late Triassic to Early Jurassic British fissure deposits. Like some trilophosaurs, it has an edentulous, or toothless beak. Tricuspisaurus gets its name from its heterodont dentition, which includes tricuspid teeth, or teeth with three cusps. The type species, T. thomasi, was named in 1957 along with the possible trilophosaur Variodens inopinatus from Somerset, England.
Daemonosaurus is an extinct genus of possible theropod dinosaur from the Late Triassic of New Mexico. The only known fossil is a skull and neck fragments from deposits of the latest Triassic Chinle Formation at Ghost Ranch. Daemonosaurus was an unusual dinosaur with a short skull and large, fang-like teeth. It lived alongside early neotheropods such as Coelophysis, which would have been among the most common dinosaurs by the end of the Triassic. However, Daemonosaurus retains several plesiomorphic ("primitive") traits of the snout, and it likely lies outside the clade Neotheropoda. It may be considered a late-surviving basal theropod or non-theropod basal saurischian, possibly allied to other early predatory dinosaurs such as herrerasaurids or Tawa.
The Colorado City Formation is a Late Triassic geologic formation in the Dockum Group of Texas, United States. It has previously been known as the Iatan Member, Colorado City Member or 'Pre-Tecovas Horizon'.
Allokotosauria is a clade of early archosauromorph reptiles from the Middle to Late Triassic known from Asia, Africa, North America and Europe. Allokotosauria was first described and named when a new monophyletic grouping of specialized herbivorous archosauromorphs was recovered by Sterling J. Nesbitt, John J. Flynn, Adam C. Pritchard, J. Michael Parrish, Lovasoa Ranivoharimanana and André R. Wyss in 2015. The name Allokotosauria is derived from Greek meaning "strange reptiles" in reference to unexpected grouping of early archosauromorph with a high disparity of features typically associated with herbivory.
Nhandumirim is a genus of basal sauropodomorph dinosaur from the Carnian age of Late Triassic Brazil. It is currently considered a saturnaliid sauropodomorph. The type and only species, Nhandumirim waldsangae, is known from a single immature specimen including vertebrae, a chevron, pelvic material, and a hindlimb found in the Santa Maria Formation in Rio Grande do Sul.