Paratypothorax

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Paratypothorax
Temporal range: Late Carnian? - Rhaetian, 216–201.6  Ma
Paratypothorax andressorum.JPG
Type specimen SMNS 5721 on display
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Order: Aetosauria
Family: Stagonolepididae
Tribe: Paratypothoracini
Genus: Paratypothorax
Long and Ballew, 1985
Type species
Paratypothorax andressi
Long and Ballew, 1985
Species
  • P. andressorumLong and Ballew, 1985 emend. Heckert and Lucas, 2000 (type)

Paratypothorax is an extinct genus of aetosaur, known from a single species, Paratypothorax andressorum. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimens come from Germany, though for more than a century they were mistakenly considered phytosaur armor. Paratypothorax was a large and wide-bodied typothoracine aetosaur, as well as the namesake of the tribe Paratypothoracisini. [1]

Contents

Discovery and distribution

1894 restoration of "Belodon", based on the skull of Nicrosaurus and the carapace of the aetosaur now known as Paratypothorax Belodon.jpg
1894 restoration of " Belodon", based on the skull of Nicrosaurus and the carapace of the aetosaur now known as Paratypothorax

Paratypothorax was first known from specimens collected from the Heslach area near Stuttgart in Germany. These hail from the Stubensandstein (also known as the Löwenstein Formation), a mid-Norian age geological unit. Heslach has also produced many fossils of a smaller aetosaur, Aetosaurus . The holotype specimen of Paratypothorax, SMNS 5721, consists of a series of articulated osteoderms alongside a few hip and leg bones. Large osteoderms of Paratypothorax were long misidentified as phytosaur armor, and referred to Belodon . Among this formerly misidentified material is SMNS 19003, a fossil unearthed in 1945 from the Schlipf Quarry near Murrhardt. Further preparation revealed that SMNS 19003 was a complete skeleton with a well-preserved skull and fully articulated carapace. [1] Paratypothorax was first recognized as an aetosaur in 1953, and was named as a new genus in 1985. [2] [3]

In 1992, Paratypothorax fossils were first identified in the Chinle Group of the southwestern United States. It has been found in the Sonsela Member of Petrified Forest National Park in Arizona, as well as the Bluewater Creek Formation of western New Mexico. It is also common in strata of the Dockum Group further in eastern New Mexico and Texas. [2] [4] Osteoderms of Paratypothorax have also been reported from the Norian-age Fleming Fjord Formation in Greenland, [5] the Zarzaitine Series of Algeria, and an unverified occurrence in India. [6]

Description

Paratypothorax possesses paramedian scutes that are wide, strap-like, and have grooves and pits on them forming radial patterns. Like other typothoracisines such as Typothorax , the lateral scutes bear large horns that are posteriorly hooked. [7] The rear of each scute is overlapped by a prominent knob. [8]

Related Research Articles

Aetosaur Extinct order of heavily armoured reptiles

Aetosaurs were heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site that they are found in.

<i>Revueltosaurus</i> Extinct genus of reptiles

Revueltosaurus is an extinct genus of suchian pseudosuchian from Late Triassic deposits of New Mexico, Arizona and North Carolina, United States. Many specimens, mostly teeth, have been assigned to Revueltosaurus over the years. Currently, three species are included in this genus, all of which were originally thought to represent monospecific genera of basal ornithischian dinosaurs. It was 1 meter long.

<i>Aetosaurus</i> Extinct genus of reptiles

Aetosaurus is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: A. ferratus, the type species from Germany and Italy; A. crassicauda from Germany; and A. arcuatus from eastern North America. Additional specimens referred to Aetosaurus have been found in the Chinle Group of the southwestern United States, and the Fleming Fjord Formation of Greenland. Specimens of Aetosaurus occur in Norian-age strata.

Calyptosuchus is an extinct genus of aetosaur from the Late Triassic of North America. Like other aetosaurs, it was heavily armored and had a pig-like snout used to uproot plants.

<i>Nicrosaurus</i> Extinct genus of reptiles

Nicrosaurus (/nɛkroʊˈsɔrəs/) is an extinct genus of phytosaur reptile existing during the Late Triassic period. Although it looked like a crocodile, it was not closely related to these creatures, instead being an example of parallel evolution. The main difference between Nicrosaurus and modern crocodiles is the position of the nostrils – Nicrosaurus's nostrils, or external nares, were placed directly in front of the forehead, whereas in crocodiles, the nostrils are positioned on the end of the snout. A 2013 study has also found that ilium of Nicrosaurus is quite distinctive from all other phytosaurs.

<i>Aetosauroides</i> Extinct genus of reptiles

Aetosauroides is an extinct genus of aetosaur from the Late Triassic of South America. It is one of four aetosaurs known from South America, the others being Neoaetosauroides, Chilenosuchus and Aetobarbakinoides. Three species have been named: the type species A. scagliai, A. subsulcatus and A. inhamandensis. Fossils have been found in the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Santa Maria Formation in the Paraná Basin in southeastern Brazil. The strata date to the late Carnian and early Norian stages, making Aetosauroides one of the oldest aetosaurs.

<i>Doswellia</i> Extinct genus of reptiles

Doswellia is an extinct genus of archosauriform from the Late Triassic of North America. It is the most notable member of the family Doswelliidae, related to the proterochampsids. Doswellia was a low and heavily built carnivore which lived during the Carnian stage of the Late Triassic. It possesses many unusual features including a wide, flattened head with narrow jaws and a box-like rib cage surrounded by many rows of bony plates. The type species Doswellia kaltenbachi was named in 1980 from fossils found within the Vinita member of the Doswell Formation in Virginia. The formation, which is found in the Taylorsville Basin, is part of the larger Newark Supergroup. Doswellia is named after Doswell, the town from which much of the taxon's remains have been found. A second species, D. sixmilensis, was described in 2012 from the Bluewater Creek Formation of the Chinle Group in New Mexico; however, this species was subsequently transferred to a separate doswelliid genus, Rugarhynchos. Bonafide Doswellia kaltenbachi fossils are also known from the Chinle Formation of Arizona.

<i>Typothorax</i> Extinct genus of reptiles

Typothorax is an extinct genus of typothoracine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum.

Coahomasuchus is an extinct genus of aetosaurine aetosaur. Remains of the genus have been found from deposits in Texas and North Carolina that date to the Otischalkian faunachron of the Late Triassic. It was small for an aetosaur, being less than 1.5 metres long. The dorsal plates are distinctively flat and unflexed, and have a faint sub-parallel to radial ornamentation. The genus lacked spines or keels on these plates, features seen in many other aetosaurs. Coahomasuchus was very similar in appearance to the closely related Aetosaurus.

<i>Dyoplax</i> Extinct genus of reptiles

Dyoplax is an extinct genus of pseudosuchian archosaur, possibly an erpetosuchid. Fossils have been found from the type locality within the upper Schilfsandstein Formation in Stuttgart, Germany. The holotype specimen was a natural cast of a nearly complete skeleton that lacked only parts of the tail and limb bones.

Ebrachosaurus is an extinct genus of aetosaur. It was named after the town of Ebrach, Germany, near an outcrop of the Blasensandstein Formation where the original fossils have been found. Other Blasensandstein fauna include the temnospondyl Metoposaurus and the phytosaur Francosuchus. The genus has often been considered synonymous with the closely related Stagonolepis. The holotype specimen was lost during World War II, so its relationships within Stagonolepididae remain indeterminant.

Rioarribasuchus is a genus of aetosaur. Fossils have been found from the Chinle Formation in Arizona and New Mexico that date back to the upper Late Carnian stage of the Late Triassic.

<i>Leptosuchus</i> Extinct genus of reptiles

Leptosuchus is an extinct genus of leptosuchomorph phytosaur with a complex taxonomical history. Fossils have been found from the Dockum Group and lower Chinle Formation outcropping in Texas, New Mexico, and Arizona, USA, and date back to the Carnian stage of the Late Triassic.

Lucasuchus is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the Late Triassic. The genus was named in 1995 after the American paleontologist Spencer G. Lucas.

<i>Machaeroprosopus</i> Extinct genus of reptiles

Machaeroprosopus is an extinct genus of mystriosuchin leptosuchomorph phytosaur from the Late Triassic of the southwestern United States. M. validus, once thought to be the type species of Machaeroprosopus, was named in 1916 on the basis of three complete skulls from Chinle Formation, Arizona. The skulls have been lost since the 1950s, and a line drawing in the original 1916 description is the only visual record of the specimen. Another species, M. andersoni, was named in 1922 from New Mexico, and the species M. adamanensis, M. gregorii, M. lithodendrorum, M. tenuis, and M. zunii were named in 1930. Most species have been reassigned to the genera Smilosuchus, Rutiodon, or Phytosaurus. Until recently, M. validus was considered to be the only species that has not been reassigned. Thus, Machaeroprosopus was considered to be a nomen dubium or "doubtful name" because of the lack of diagnostic specimens that can support its distinction from other phytosaur genera. However, a taxonomic revision of Machaeroprosopus, conducted by Parker et al., revealed that UW 3807, the holotype of M. validus, is not the holotype of Machaeroprosopus, while the previously named species Pseudopalatus buceros has priority as the type species of the genus. Therefore, the name Pseudopalatus must be considered a junior synonym of Machaeroprosopus, and all species of the former must be reassigned to the later. This revised taxonomy was already accepted in several studies, including Stocker and Butler (2013). Stocker and Butler (2013) also treated M. andersoni as a valid species, and not a junior synonym of Machaeroprosopus buceros as was previously suggested by Long and Murry (1995).

Tecovasuchus is an extinct genus of aetosaur. It is known primarily from osteoderms found from the Tecovas Formation in Texas, which is Late Triassic in age, dating back to the lower Norian. Material is also known from several other localities of the Chinle Group in New Mexico and Arizona, such as older Carnian outcrops and younger Rhaetian outcrops. Specimens of Tecovasuchus have been collected from the Tecovas Formation, the Bluewater Creek Formation, and the Los Esteros Member of the Santa Rosa Formation.

Aetosaurinae Extinct subfamily of reptiles

Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.

Redondasuchus is an extinct genus of aetosaur. It may be a junior synonym of Typothorax coccinarum, another aetosaur. Redondasuchus is a member of the clade Typothoracisinae within the subfamily Aetosaurinae, and lived during the middle Norian stage of the Late Triassic. Material belonging to the genus has been found from the Redonda Formation in east-central New Mexico. The type species, R. reseri, was named in 1991 after having been referred to as a species of Typothorax since 1985. A second species, R. rineharti, was described in 2006.

Gorgetosuchus is an extinct genus of aetosaur from the Late Triassic of the North Carolina, represented by the type species Gorgetosuchus pekinensis. It is mainly known from osteoderms, including the front half of an articulated carapace. Gorgotesuchus is typically considered a basal desmatosuchin, though alternative interpretations exist.

Apachesuchus is an extinct genus of aetosaur from the Late Triassic of New Mexico. It is only known from several paramedian osteoderms discovered in Quay County in eastern New Mexico. This area belongs to the late Norian-age Quay Member of the Redonda Formation. Unique among aetosaurs, its osteoderms are nearly completely smooth, without strong pits or grooves. The left dorsal paramedian has a relatively high width-to-length ration, suggesting that Apachesuchus is a wide-bodied aetosaur within the clade Typothoracinae.

References

  1. 1 2 Schoch, Rainer R.; Desojo, Julia Brenda (2016). "Cranial anatomy of the aetosaur Paratypothorax andressorum Long & Ballew, 1985, from the Upper Triassic of Germany and its bearing on aetosaur phylogeny". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 279 (1). doi:10.1127/njgpa/2016/0542. ISSN   0077-7749.
  2. 1 2 Long, R. A.; Ballew, K. L. (1985). "Aetosaur dermal armor from the Late Triassic of southwestern North America, with special reference to material from the Chinle Formation of Petrified Forest National Park". Museum of Northern Arizona Bulletin. 47: 45–68.
  3. Parrish, J. M. (1994). "Cranial osteology of Longosuchus meadei and the phylogeny and distribution of the Aetosauria". Journal of Vertebrate Paleontology. 14 (2): 196–209. doi:10.1080/02724634.1994.10011552.
  4. Hunt, A. P.; Lucas, S. G. (1992). "The first occurrence of the aetosaur Paratypothorax andressi (Reptilia:Aetosauria) in the western United States and its biochronological significance". Paläontologische Zeitschrift. 66 (1–2): 147–157. doi:10.1007/BF02989485. S2CID   129434871.
  5. Jenkins, F. A. Jr.; Shubin, N. H.; Amaral, W. W.; Gatesy, S. M.; Schaff, C. R.; Clemmensen, L. B.; Downs, W. R.; Davidson, A. R.; Bonde, N.; Osbaeck, F. (1994). "Late Triassic continental vertebrates and depositional environments of the Fleming Fjord Formation, Jameson Land, East Greenland". Meddelelser om Grønland, Geoscience. 32: 1–25.
  6. Lucas, Spencer G.; Heckert, Andrew B.; Rinehart, Larry F. (2006). "The Late Triassic aetosaur Paratypothorax" (PDF). New Mexico Museum of Natural History and Science Bulletin. 37: 575–580.
  7. Murry, P. A. (1986). "Vertebrate paleontology of the Dockum Group". In K. Padian (ed.). The Beginning of the Age of Dinosaurs. Cambridge: Cambridge University Press. pp. 109–138.
  8. Heckert, A. B.; Lucas, S. G. (1999). "A new aetosaur (Reptilia: Archosauria) from the Upper Triassic of Texas and the phylogeny of aetosaurs". Journal of Vertebrate Paleontology. 19 (1): 50–68. CiteSeerX   10.1.1.563.9516 . doi:10.1080/02724634.1999.10011122.