Rioarribasuchus Temporal range: | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Aetosauria |
Family: | † Stagonolepididae |
Genus: | † Rioarribasuchus Lucas, Hunt & Spielmann, 2006 |
Species: | †R. chamaensis |
Binomial name | |
†Rioarribasuchus chamaensis (Zeigler, Heckert & Lucas, 2003) | |
Synonyms | |
|
Rioarribasuchus is a genus of aetosaur. Fossils have been found from the Chinle Formation in Arizona and New Mexico that date back to the upper Late Carnian stage of the Late Triassic.
"Desmatosuchus" chamaensis was named in 2003 and found from the Petrified Forest Member of the Chinle Formation in New Mexico. [1] [2] It was suggested to be more closely related to Paratypothorax , and so Parker gave it the name Heliocanthus. However, this new generic name was first proposed in an unpublished thesis, and thus did not meet ICZN regulations for the naming of a new taxon.
Later published papers reasserted the genetic separation of "D". chamaensis from Desmatosuchus, but the name Heliocanthus remained a nomen nudum until 2007, where it was thoroughly rediscribed in a paper published by the Journal of Systematic Palaeontology . [3]
However, a paper previously published in late 2006 assigned "D". chamaensis to the new genus Rioarribasuchus. [4] As a result, Heliocanthus is a junior objective synonym of Rioarribasuchus because the genus has seniority over Heliocanthus. However, the name Rioarribasuchus has been viewed as a violation of the code of ethics laid out in Appendix A of the International Code of Zoological Nomenclature and the papers that made use of the name have even been described as practicing "intellectual theft". [5]
An article published later in 2007 in the science blog Tetrapod Zoology brought these events to the attention of a wider range of readers, and the controversy was dubbed "Aetogate". [6] This sparked continued debate regarding these issues among vertebrate paleontologists, which eventually led to an investigation by the Society of Vertebrate Paleontology into these issues and a response given in mid 2008 regarding the unethical conduct of the authors who described Rioarribasuchus. [7]
Aetosaurs were heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site that they are found in.
Aetosaurus is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: A. ferratus, the type species from Germany and Italy; A. crassicauda from Germany; and A. arcuatus from eastern North America. Additional specimens referred to Aetosaurus have been found in the Chinle Group of the southwestern United States, and the Fleming Fjord Formation of Greenland. Specimens of Aetosaurus occur in Norian-age strata.
Stagonolepis is an extinct genus of stagonolepidid aetosaur known from the Late Triassic Hassberge Formation of Germany, the Drawno Beds of Poland, the Lossiemouth Sandstone of Scotland, the Chinle Formation of Arizona and Utah and the Bluewater Creek Formation of New Mexico.
Desmatosuchus is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic.
Calyptosuchus is an extinct genus of aetosaur from the Late Triassic of North America. Like other aetosaurs, it was heavily armored and had a pig-like snout used to uproot plants.
Aetosauroides is an extinct genus of aetosaur from the Late Triassic of South America. It is one of four aetosaurs known from South America, the others being Neoaetosauroides, Chilenosuchus and Aetobarbakinoides. Three species have been named: the type species A. scagliai, A. subsulcatus and A. inhamandensis. Fossils have been found in the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Santa Maria Formation in the Paraná Basin in southeastern Brazil. The strata date to the late Carnian and early Norian stages, making Aetosauroides one of the oldest aetosaurs.
Typothorax is an extinct genus of typothoracine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum.
Coahomasuchus is an extinct genus of aetosaurine aetosaur. Remains of the genus have been found from deposits in Texas and North Carolina that date to the Otischalkian faunachron of the Late Triassic. It was small for an aetosaur, being less than 1.5 metres long. The dorsal plates are distinctively flat and unflexed, and have a faint sub-parallel to radial ornamentation. The genus lacked spines or keels on these plates, features seen in many other aetosaurs. Coahomasuchus was very similar in appearance to the closely related Aetosaurus.
Ebrachosaurus is an extinct genus of aetosaur. It was named after the town of Ebrach, Germany, near an outcrop of the Blasensandstein Formation where the original fossils have been found. Other Blasensandstein fauna include the temnospondyl Metoposaurus and the phytosaur Francosuchus. The genus has often been considered synonymous with the closely related Stagonolepis. The holotype specimen was lost during World War II, so its relationships within Stagonolepididae remain indeterminant.
Lucasuchus is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the Late Triassic. The genus was named in 1995 after the American paleontologist Spencer G. Lucas.
Paratypothorax is an extinct genus of aetosaur, known from a single species, Paratypothorax andressi. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimens come from Germany, though for more than a century they were mistakenly considered phytosaur armor. Paratypothorax was a large and wide-bodied typothoracine aetosaur, as well as the namesake of the tribe Paratypothoracisini.
Tecovasuchus is an extinct genus of aetosaur. It is known primarily from osteoderms found from the Tecovas Formation in Texas, which is Late Triassic in age, dating back to the lower Norian. Material is also known from several other localities of the Chinle Group in New Mexico and Arizona, such as older Carnian outcrops and younger Rhaetian outcrops. Specimens of Tecovasuchus have been collected from the Tecovas Formation, the Bluewater Creek Formation, and the Los Esteros Member of the Santa Rosa Formation.
Typothoracinae is a clade of aetosaurs within the subfamily Aetosaurinae. It was originally defined as a stem-based taxon including all aetosaurs closer to Typothorax than to Stagonolepis or Desmatosuchus. This definition was later expanded to specifically exclude Aetosaurus; as of 2016, Typothoracinae is defined as the least inclusive clade containing Typothorax and Paratypothorax, but not Aetosaurus,Stagonolepis, or Desmatosuchus. The clade was first named in 2007 under the spelling Typothoracisinae, after its namesake Typothorax. However, this spelling was based on incorrect taxonomic nomenclature, and the clade's name was corrected to Typothoracinae in 2016.
Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.
Paratypothoracini is a clade of aetosaurs within the group Typothoracinae. It is a node-based taxon that includes Rioarribasuchus (=Heliocanthus), Paratypothorax, Tecovasuchus, and all descendants of their most recent common ancestor. The clade was first named in 2007 under the spelling Paraypothoracisini, after its namesake Paratypothorax. However, this spelling was based on incorrect taxonomic nomenclature, and the clade's name was corrected to Paratypothoracinae in 2016.
Desmatosuchinae is a major subfamily of aetosaurs within the clade Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Desmatosuchus than to Stagonolepis,Aetosaurus, or Paratypothorax.
Redondasuchus is an extinct genus of aetosaur. It may a junior synonym of Typothorax coccinarum, another aetosaur. Redondasuchus is a member of the clade Typothoracisinae within the subfamily Aetosaurinae, and lived during the middle Norian stage of the Late Triassic. Material belonging to the genus has been found from the Redonda Formation in east-central New Mexico. The type species, R. reseri, was named in 1991 after having been referred to as a species of Typothorax since 1985. A second species, R. rineharti, was described in 2006.
The Bluewater Creek Formation is a geologic formation in west-central New Mexico. It preserves fossils dating back to the late Triassic period.
Apachesuchus is an extinct genus of aetosaur from the Late Triassic of New Mexico. It is only known from several paramedian osteoderms discovered in Quay County in eastern New Mexico. This area belongs to the late Norian-age Quay Member of the Redonda Formation. Unique among aetosaurs, its osteoderms are nearly completely smooth, without strong pits or grooves. The left dorsal paramedian has a relatively high width-to-length ration, suggesting that Apachesuchus is a wide-bodied aetosaur within the clade Typothoracinae.
{{cite web}}
: CS1 maint: archived copy as title (link){{cite web}}
: CS1 maint: archived copy as title (link)