Agnatha is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes.
Heterostraci is an extinct subclass of pteraspidomorph, Ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian.
Thelodonti is a class of extinct Palaeozoic jawless fishes with distinctive scales instead of large plates of armor.
Anaspida is an extinct group of jawless fish that existed from the early Silurian period to the late Devonian period. They were classically regarded as the ancestors of lampreys, but it is denied in recent phylogenetic analysis, although some analysis show these group would be at least related. Anaspids were small marine fish that lacked a heavy bony shield and paired fins, but were distinctively hypocercal.
Galeaspida is an extinct taxon of jawless marine and freshwater fish. The name is derived from galea, the Latin word for helmet, and refers to their massive bone shield on the head. Galeaspida lived in shallow, fresh water and marine environments during the Silurian and Devonian times in what is now Southern China, Tibet and Vietnam. Superficially, their morphology appears more similar to that of Heterostraci than Osteostraci, there being currently no evidence that the galeaspids had paired fins. A galeaspid Tujiaaspis vividus from the Silurian period of China was described in 2022 as having a precursor condition to the form of paired fins seen in Osteostraci and gnathostomes. Earlier than this, Galeaspida were already in fact regarded as being more closely related to Osteostraci, based on the closer similarity of the morphology of the braincase.
Thelodus is an extinct genus of thelodont agnathan that lived during the Silurian period. Fossils have been found in Europe, Asia and North America. Unlike many thelodonts, species of Thelodus are known not only from scales, but from impressions in rocks. Some species, such as the Canadian T. inauditus, are thought to be comparable in size to other thelodonts, i.e., from 5 to 15 centimeters in length. The scales of the type species, T. parvidens of Silurian Great Britain, however, reach the size of coins, and, if proportioned like other thelodonts, such as Loganellia, the living animal would have been about one meter in length.
Jamoytius kerwoodi is an extinct species of primitive, eel-like jawless fish known from the Patrick Burn Formation in Scotland, dating to the Llandovery epoch of the Early Silurian period.
Phlebolepis is an extinct thelodont agnathan genus belonging to the family Phlebolepididae. Whole fossils are found in Late Silurian aged strata from Saaremaa, Estonia. Phlebolepis elegans was average-sized for a thelodont, 7 cm long.
Furcacauda is a genus of thelodontid agnathan from the Lower Devonian of Canada, and is the type genus of the order Furcacaudiformes. Furcacaudiform thelodontids were deep water jawless vertebrates with symmetrical fork and lobed-finned tails and scales smaller than typical loganellid and nikoliviid thelodonti scales. Furcacaudiform thelodonts are noted as having a laterally compressed body, large anterior eyes, slightly posterior, lateral, and vertical to a small mouth, and a condensed curved row of branchial openings (gills) directly posterior to the eyes. Many but not all had laterally paired fins. Wilson and Caldwell also note the presence of a caudal peduncle and a long caudal fin made of two large lobes, one dorsal and one ventral separated by 8 to 14 smaller intermediate lobes, giving the appearance of a striated half-moon shaped tail resembling the tail of a heterostracan. A large square cavity within the gut connecting a small intestine to an anal opening lead many to believe that it is this genus that exhibits the first vertebrate stomach. According to Wilson and Caldwell their discovery, based on sediment infillings of fossils of the Furcacauda heintze, gives credence to the evolutionary development of stomach before jaws.
Sphenonectris is an extinct genus of thelodont, which lived in Canada during the Early Devonian period.
Cometicercus is an extinct genus of thelodont which lived in Canada during the Early Devonian period.
Longodus is an extinct genus of thelodont, placed in its own family – Longodidae – which existed in what is now Estonia during the Ludlow epoch of the upper Silurian period. The type and only species is Longodus acicularis. They are most noted for their long, needle-like scales, which run vertically along their trunk.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada.
Furcacaudiformes is an extinct order of jawless fish in the class Thelodonti.
Thelodontiformes is an extinct order of jawless fish of the Silurian.
Archipelepidiformes is an order of extinct jawless fishes in the class Thelodonti.
Archipelepis is a genus of extinct thelodont agnathans, and are the most primitive recognized thelodonts of which whole body fossils are known. Fossils of bodies and scales are currently known from Late Telychian to Wenlock-aged marine strata of northern Canada.
Phlebolepididae is an extinct thelodont agnathan family in the order Phlebolepidiformes.
Erepsilepis is an extinct thelodont agnathan genus in the family Phlebolepididae.
