The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae. Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 45 millimeters, with 55 to 75 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus easoni is a species of soil centipede in the family Geophilidae found throughout Britain and Ireland, though its range extends through western France to at least the foothills of the Pyrenees. Until 2001, it was considered synonymous with G. carpophagus, and most pre-2001 records probably refer to G. easoni. It's typically shorter than G. carpophagus, with fewer leg pairs, uniform tan/chestnut coloring, and a greater size and number of coxal pores, as well as a darkly pigmented mid-piece of the labrum which bears blunt teeth. Specimens in north-western Iberia typically have a wider range of leg pairs. It nests on the ground surface under stones and dead wood.
Geophilus aenariensis is a species of soil centipede in the family Geophilidae found in the southern Italian Peninsula. It has a body length of up to 22 millimeters, around 47 leg pairs in females and 51 in males, an absence of anal pores and bristles on the lateral part of the labrum, a distinct carpophagus pit, a small but pointed pretarsus of the second maxillae, and 3-5 coxal pores in each coxopleuron with no isolated coxal pores.
Geophilus mordax, also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 millimeters in length, though it averages 25–40, has 49–53 leg pairs in males and 49–57 in females, and is bright red in color. G. mordax also bears 3–5 labral teeth, a short and robust apical claw of the second maxillae, an exposed prebasal plate, and a large, consolidated sacculi.
Geophilus nanus is a species of soil centipede in the family Geophilidae found on the Iberian Peninsula. It's a poorly defined species described as being pale yellow, 15 mm long, with smooth sternites bearing no furrows, final hip with few pores next to the sternite rim, tergites deeply double-furrowed from the basal shield onward, and 41 leg pairs. It’s apparently related to G. gracilis, differentiated by jaws that do not surpass the forehead or bear chitin lines, and by the 2nd tarsal segment of the final leg pair being just a tiny stub.
Geophilus anonyx is a species of soil centipede in the family Geophilidae found in Oregon. It was originally placed in the genus Brachygeophilus based on the lack of sternal pores, a character shared with the type species B. truncorum, however it was later moved to Geophilus.
Geophilus nicolanus is a species of soil centipede in the family Geophilidae found on San Nicholas Island, California. It's dark brown in color and grows up to 62 millimeters long, with 63 leg pairs, a concealed prebasal plate, numerous coxal pores, and a very wide final ventral plate with convex sides.
Geophilus richardi is a species of soil centipede in the family Geophilidae found in France, Italy, and the Ionian Islands. Females of this species have 33 pairs of legs; males have 29 or 31. This species is one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs. This species grows up to 10 millimeters long, has no carpophagus pit or pore-fields, and has a gradually tapering, curved pretarsus of the second maxillae. G. richardi lacks typical ventral pores between 2–4mm. The sternites instead bear a small number of pores between 0.5–1mm that differ from micropores, which are unbounded by a cuticular ring. These are possibly the remnants of typical ventral pores, their smaller size being a byproduct of overall miniaturization.
Geophilus pygmaeus is a species of soil centipede in the family Geophilidae found in the Southern Limestone Alps, between the Bergamasque Prealps in the west and the Slovene Prealps in the east, and in the northernmost Dinarides. Past records from other areas are usually false. G. pygmaeus is less than 20 millimeters long and has 41-47 leg pairs. It differs from other European species of Geophilus by the shape of the forcipular apparatus and arrangement of coxal pores.
Geophilus tampophor is a species of soil centipede in the family Geophilidae found in Oregon. It was originally placed in the genus Brachygeophilus, however it was later moved to Geophilus. Like G. anonyx, it differs from other western American geophiliids by having prehensorial claws armed at the base, however unlike G. anonyx it possesses armed anal claws and a lesser number of legs.
Geophilus bipartitus is a species of soil centipede in the family Geophilidae found in Japan. It grows up to 15 millimeters in length; the males have about 35 leg pairs, the females 39. It lives in Japanese white birch.
Geophilus rhomboideus is a species of soil centipede in the family Geophilidae found in Japan. It grows up to 30 millimeters in length; the males have about 43 leg pairs, the females 49.
Geophilus sounkyoensis is a species of soil centipede in the family Geophilidae found in Japan. Its body is yellow in color and it grows up to 40 millimeters in length; the males have about 55 leg pairs, the females 57.
Geophilus bluncki is a species of soil centipede in the family Geophilidae found in San Remo, Italy. It grows up to 23 millimeters in length; the males have about 61 leg pairs. The uniform pore fields and long antennae resemble Arctogeophilus glacialis, formerly Geophilus glacialis.
Geophilus monoporus is a species of soil centipede in the family Geophilidae found in Tiba, Japan. It grows up to 18 millimeters in length; it's named for the single pore at the base of the final leg pair.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 37 to 41 pairs of legs; females have 35 to 41.
Geophilus koreanus is a species of soil centipede in the family Geophilidae found in North Korea. It's yellow in color and grows up to 30 millimeters long, with 69 leg pairs, a clypeus rather longer than wide, filiform antennae, central part of the labrum with 8 teeth, maxilla completely fused without median suture, tergite bifurcate, final leg tarsus bipartite, and a clawed pratarsus. It's similar to G. strictus, though the latter differs by the middle part of the labrum bearing a large number of very small teeth, and the final hip bearing a large number of pores.
Geophilus bosniensis is a species of soil centipede in the family Geophilidae endemic to Bosnia and Herzegovina. It grows up to 30 millimeters and has 75 leg pairs, as well as sternites unseparated in the median but with a suture line, and sternal pore areas in the trunk segments only. Overall, the identity and phyletic position of this centipede are uncertain.
