Geophilidae | |
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Geophilus flavus , the Netherlands | |
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Family: | Geophilidae |
Synonyms [2] | |
Chilenophilidae |
Geophilidae is a family of soil centipedes in the superfamily Geophiloidea and the order Geophilomorpha. [3] [4] In 2014, a phylogenetic analysis based on morphological and molecular data found this family to be polyphyletic. [3] To avoid this polyphyly, authorities dismissed the families Aphilodontidae, [5] Dignathodontidae, [6] Linotaeniidae, [7] and Macronicophilidae, [8] which are now deemed to be junior synonyms for Geophilidae. [4] Authorities also moved some genera from Geophilidae to form the family Zelanophildae in order to avoid the polyphyly of the family Geophilidae. [3] [9] The family Geophilidae now includes more than 650 species in more than 120 genera. [4] This family has a cosmopolitan distribution, with species found almost worldwide. [10]
Species in this family are characterized by mandibles with a single pectinate lamella. Sternal pores are often present but with variable arrangements, and coxal organs are usually present and open into pits or through distinct pores. Female gonopods in this family are usually an undivided lamina. [10]
Compared to most other families in the suborder Adesmata, this family features a modest number of leg-bearing segments and limited variation in this number within each species. [11] Two European species in this family include centipedes with only 29 pairs of legs: Geophilus persephones (29 in the only specimen, a male), and G. richardi (29 or 31 in males and 33 in females). [12] [13] In the order Geophilomorpha, only two species include centipedes with fewer leg pairs, both of them in the family Schendylidae. [12] Several other species in the family Geophilidae are known from specimens with notably few leg pairs in each sex, including Ribautia platensis (as few as 31 in each sex), [14] G. hadesi (33 in each sex), [15] Schendyloides alacer (as few as 33 in each sex), [16] and Strigamia sibirica (as few as 33 in each sex). [17]
This family contains these genera: [1]
Geophilomorpha is an order of centipedes commonly known as soil centipedes. The name "Geophilomorpha" is from Ancient Greek roots meaning "formed to love the earth." This group is the most diverse centipede order, with 230 genera. These centipedes are found nearly worldwide but are absent in Antarctica and most Arctic regions.
Pachymerium caucasicum is a species of centipede in Geophilidae family. It was described by Carl Attems in 1903 and is endemic to the European part of Turkey. Males of this species have 47 pairs of legs; females have 49 pairs of legs. Authorities now deem P. caucasicum to be a junior synonym of P. ferrugineum.
Pachymerium tabacarui is a species of centipede in Geophilidae family that is endemic to Romania. The original description of this species is based on a female specimen from the Carpathians measuring 15 mm in length with 53 pairs of legs. Some authorities consider this description consistent with a juvenile specimen of P. ferrugineum and therefore deem P. tabacarui to be a junior synonym of that species. A more recent review of this description suggests that P. tabacarui is a junior synonym of Geophilus flavus rather than P. ferrugineum. Extensive investigations in the Carpathian region have failed to collect any more specimens.
Geophilus hadesi is a species of soil centipede in the family Geophilidae. This centipede is a troglobite, spending its entire life cycle in a cave environment. This species and Geophilus persephones are the only two troglobites known in the order Geophilomorpha. The species G. hadesi is named after Hades, god of the underworld in Greek mythology and the husband of Persephone, the namesake of the first troglobite discovered among the soil centipedes. The species G. hadesi has been observed in a cave as far as 1,100 meters below the surface, the deepest underground that any centipede has ever been recorded.
Mecistocephalidae is a monophyletic family of centipedes in the order Geophilomorpha. It is the only family in the suborder Placodesmata. Most species in this family live in tropical or subtropical regions, but some occur in temperate regions. This family is the third most diverse in the order Geophiliomorpha, with about 170 species, including about 130 species in the genus Mecistocephalus.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus. This genus has a Holarctic distribution.
Linotaeniidae are a monophyletic clade of soil centipedes in the family Geophilidae found mostly in the temperate regions of the Holarctic as well as the south Andes. Species in the clade Linotaeniidae are characterized by a body that usually tapers toward the anterior tip; mandibles with a single pectinate lamella; second maxillae with coxo-sternite usually undivided and claws without projections; forcipular segment short, with tergite remarkably wide, forcipules evidently tapering; coxal organs opening through distinct pores on the ventral surface of the coxo-pleura. The number of legs in this clade varies within as well as among species and ranges from as few as 31 pairs of legs to as many as 83 leg pairs. Compared to most families in the suborder Adesmata, this clade features a modest number of leg-bearing segments and limited variation in this number within each species.
Schendylidae is a family of soil centipedes in the superfamily Himantarioidea and the order Geophilomorpha. These centipedes are found in the Americas, the Palearctic region, Africa, Madagascar, Australia, and southeast Asia, and also on some Pacific islands.
Geophilus persephones is a species of soil centipede in the family Geophilidae discovered in 1999. This species is named after Persephone, the queen of the underworld in Greek mythology, and found in caves in the Gouffre de la Pierre Saint-Martin. It has elongated antennae and legs as well as abundant sensory setae, and like other geophilomorhps it lacks sight, has a flattened trunk, and is well adapted to underground life. This species was the first troglomorphic geophilomorph ever discovered and one of the only two in existence along with Geophilus hadesi. Known from a single male specimen measuring 16.2 mm in length, G. persephones has only 29 pairs of legs, one of only two species in the Geophilidae family to feature so few leg pairs.
Acanthogeophilus spiniger is a species of soil centipede in the family Geophilidae found in Northwest Africa. The original description of this species is based on a male specimen measuring 27 mm in length with 71 pairs of legs. It was first assigned to the genus Geophilus, but was moved to Acanthogeophilus in 1999 by Foddai and Minelli. Like other species in its genus, it is characterized by incomplete chitin lines, complete coxopleural sutures, stout legs, and a claw-like pretarsus.
Geophilus arenarius is a species of soil centipede in the family Geophilidae. This centipede is found in Algeria. As part of the carpophagus species-complex, this species is closely related to both G. carpophagus and G. easoni.
Geophilus aetnensis is a species of soil centipede in the family Geophilidae found in Europe and northern Asia, excluding China. As described by Verhoeff in 1928, it grows up to 28 millimeters and has 53 leg pairs, 4 sensory setae each on the 2nd-4th front sternites, slightly notched maxillae, and very faint sternal pits on the thorax as well as poorly developed sternal grooves.
Geophilus richardi is a species of soil centipede in the family Geophilidae. This centipede is found in France and Monaco in the Western Alps and on the Italian mainland, Sicily, Sardinia, and the Ionian Islands. This species is notable as one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs.
Geophilus pusillus is a species of soil centipede in the family Geophilidae found in Algeria. It grows up to 11 millimeters in length. The original description of this species is based on ten specimens: five male specimens from Algeria with 31 pairs of legs, and five specimens from Germany. Records from the Alpstein mountains indicate that G. pusillus is a soil-dwelling species that prefers humus-rich soil, but these records deserve confirmation. Some authorities consider the specimens recorded from Europe to be specimens of G. ribauti that have been misidentified as specimens of the Algerian species G. pusillus.
Geophilus bobolianus is a species of soil centipede in the family Geophilidae found in France and Italy. This species has 45 to 51 pairs of legs. It was originally classified as a subspecies of G. longicornis identified by its lack of anterior sternal pores.
Strigamia crassipes is a centipede belonging to the family Linotaeniidae in the order Geophilomorpha.
Strigamia acuminata, commonly called the shorter red centipede, is a centipede in the family Linotaeniidae in the order Geophilomorpha.
Geophilus oligopus is a species of soil centipede in the family Geophilidae. This centipede is found in several European countries, including Austria, the Czech Republic, Italy, Croatia, Slovenia, Bosnia and Herzegovenia, and Romania. Although this centipede has been described as having an Alpine-Dinaric distribution, this species has also been found in the Carpathian mountains and may be more widespread than previously thought.
Mecophilus tupiniquim is a species of soil centipede in the subfamily Aphilodontinae, a clade formerly known as the family Aphilodontidae but now deemed a subfamily within the family Geophilidae. This centipede is notable for its small size and modest number of legs. This species is found in Brazil.