Pachymerium

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Pachymerium
Pachymerium ferrugineum 03.JPG
Pachymerium ferrugineum
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Myriapoda
Class: Chilopoda
Order: Geophilomorpha
Family: Geophilidae
Genus: Pachymerium
C. L. Koch, 1847
Type species
Geophilus ferrugineum
C.L. Koch, 1835
Synonyms
  • Pachymerium (Eupachymerium)Attems, 1909
  • Geophilus (Khroumiriophilus)Attems, 1908

Pachymerium is a genus of soil centipedes in the family Geophilidae. [1] This genus contains more than 20 species. [2] These centipedes are found mainly in the west Palearctic region and in south Africa. [3]

Contents

Description

Centipedes in this genus feature an elongated head with an intermediate part of the labrum separating the side pieces. Each of the second maxillae ends in a claw. The forcipular tergite is distinctly narrower than the following tergite. The posterior part of the forcipular coxosternite is broad. The first article of the forcipule is elongated and features a distal denticle, and the ultimate article features a prominent basal denticle. Fields of pores appear on at least the anterior sternites, with pores in a pair of anterior groups and a posterior transverse band. The sternite of the last leg-bearing segment is either about as long as wide or longer. The basal element of each of the ultimate legs features scattered pores, and each of these legs ends in a claw. [4] [3] [5]

These centipedes range from about 2 cm to about 8 cm in length and have 37 to 79 pairs of legs. [3] The Chilean species Pachymerium armatum measures only 20 mm in length and has only 37 leg pairs, [6] the minimum number recorded in this genus. [3] The Russian species P. minutum is also notable for its small size (only 17 mm in length) and its modest number of legs (39 pairs in each sex). [7] [8] The Portuguese species P. coiffaiti is notable for its large size (males reaching 58 mm in length, females ranging from 70 mm to 85 mm) and many legs (males with 69 pairs, females with 73 or 75). [9] [8] The Cuban species P. cubanum and the large Turkish species P. serratum (measuring 50 mm and 78 mm in length, respectively) each can have as many as 79 leg pairs, [10] [11] the maximum number recorded in this genus. [3]

Phylogeny

A phylogenetic analysis of the order Geophilomorpha using both molecular data and morphology places a representative of the genus Pachymerium ( P. ferrugineum ) in a clade with a representative of Schendyloides , another genus in the family Geophilidae. This phylogenetic tree suggests that the genera Pachymerium and Schendyloides are more closely related to one another than any other genera included in this analysis. These two close relatives form a sister group for another clade formed by representatives of two more genera in the same family, Arctogeophilus and Alloschizotaenia , which emerge as the next closest relatives included in this analysis. [12]

Species

This genus contains the following species: [13]

References

  1. Bonato, L.; Chagas Junior, A.; Edgecombe, G.D.; Lewis, J.G.E.; Minelli, A.; Pereira, L.A.; Shelley, R.M.; Stoev, P.; Zapparoli, M. (2016). "Pachymerium C.L. Koch, 1847". ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Retrieved 2025-11-28.
  2. "ITIS - Report: Pachymerium". itis.gov. Retrieved 2025-11-28.
  3. 1 2 3 4 5 Bonato, Lucio; Edgecombe, Gregory D.; Zapparoli, Marzio (2011). "Chilopoda – Taxonomic overview". In Minelli, Alessandro (ed.). The Myriapoda. Volume 1. Leiden: Brill. pp. 363–443 [420]. ISBN   978-90-04-18826-6. OCLC   812207443.
  4. Dyachkov, Yurii V.; Bonato, Lucio (2024-04-23). "An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia". ZooKeys (1198): 17–54 [27]. Bibcode:2024ZooK.1198...17D. doi: 10.3897/zookeys.1198.119781 . ISSN   1313-2970. PMC   11061560 . PMID   38693975.
  5. Bonato, Lucio; Edgecombe, Gregory; Lewis, John; Minelli, Alessandro; Pereira, Luis; Shelley, Rowland; Zapparoli, Marzio (2010-11-18). "A common terminology for the external anatomy of centipedes (Chilopoda)". ZooKeys (69): 17–51. Bibcode:2010ZooK...69...17B. doi: 10.3897/zookeys.69.737 . ISSN   1313-2970. PMC   3088443 . PMID   21594038.
  6. Attems, Carl (1929). Attems, Karl (ed.). Lfg. 52 Myriapoda, 1: Geophilomorpha (in German). De Gruyter. p. 249. doi:10.1515/9783111430638.
  7. Sseliwanoff, A. (1884). "Materialii k isyceniyn russkih tyisiachenogich (Myriapoda)". Horae Societatis Entomologicae Rossicae (in Russian). 18: 69–121 [89–90] via Biodiversity Heritage Library.
  8. 1 2 Bonato, Lucio; Minelli, Alessandro (2014-03-03). "Chilopoda Geophilomorpha of Europe: a revised list of species, with taxonomic and nomenclatorial notes" . Zootaxa. 3770 (1): 1–136 [49-50, 52]. Bibcode:2014Zoot.37700.1.1B. doi:10.11646/zootaxa.3770.1.1. ISSN   1175-5334.
  9. Demange, J.M. (1959). "Myriapodes Chilopodes de Madère (Mission de M. le Pr. A. Vandel)". Revue Française d'Entomologie (in French). 26: 157–166 [162–164].
  10. Matic, Z.; Negrea, St.; Fundora Martínez, C. (1977). "Recherches sur les Chilopodes hypogés de Cuba. II". Résultats des Expéditions Biospéléologiques Cubano-Roumaines à Cuba (in French). Vol. 2. Editura Academiei Republicii Socialiste România. București. pp. 277–301 [288].
  11. Verhoeff, Karl W. (1943). "Über Chilopoden der Türkei. III. Aufsatz". Zoologischer Anzeiger (in German). 143: 116–140 [133–134].
  12. Bonato, Lucio; Drago, Leandro; Murienne, Jérôme (2014). "Phylogeny of Geophilomorpha (Chilopoda) inferred from new morphological and molecular evidence". Cladistics. 30 (5): 485–507 [498]. Bibcode:2014Cladi..30..485B. doi:10.1111/cla.12060. ISSN   0748-3007. PMID   34794246.
  13. "Pachymerium". Global Myriapod Information System. Zoologische Staatssammlung München. June 26, 2008. Archived from the original on October 11, 2013. Retrieved October 5, 2013.