The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae. Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus easoni is a species of soil centipede in the family Geophilidae found throughout Britain and Ireland, though its range extends through western France to at least the foothills of the Pyrenees. Until 2001, it was considered synonymous with G. carpophagus, and most pre-2001 records probably refer to G. easoni. It's typically shorter than G. carpophagus, with fewer leg pairs, uniform tan/chestnut coloring, and a greater size and number of coxal pores, as well as a darkly pigmented mid-piece of the labrum which bears blunt teeth. Specimens in north-western Iberia typically have a wider range of leg pairs. It nests on the ground surface under stones and dead wood.
Geophilus arenarius is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. This species is frequently misidentified with G. electricus, and as part of the carpophagus species-complex, is closely related to both G. carpophagus and G. easoni, though it differs mainly by lacking a transverse suture on the head and peculiar integumental features (carpophagus-structures) along the trunk, as well as having relatively stouter antennae and forcipular coxosternite. G. arenarius is distinctly smaller at full growth than G. carpophagus, with usually blunter and more sclerotised tubercles lining the intermediate part of the labrum and a minute denticle at the basis of the forcipular tarsungula. It has fewer bristles lining the lateral parts of the labrum than G. easoni as well as a generally higher number of legs and a more greyish coloured trunk. An examination of 36 G. arenarius syntypes indicates that males of this species have 55 pairs of legs, whereas females have 55 to 59 leg pairs, with 57 as the most common number.
Geophilus proximus is a species of soil centipede in the family Geophilidae found in the northern part of the Palearctic and widespread across the entire Baltic basin, though it reaches as far as the Arctic Circle and has been introduced through human agency to northern, central, and eastern parts of Kazakhstan. It was recorded once with certainty in Britain from Unst in the Shetland Islands; distribution in the rest of Europe is difficult to assess because of frequent misidentifications of the species. Populations from northern Europe are mostly parthenogenetic.
Geophilus aenariensis is a species of soil centipede in the family Geophilidae found in the southern Italian Peninsula. It has a body length of up to 22 millimeters, around 47 leg pairs in females and 51 in males, an absence of anal pores and bristles on the lateral part of the labrum, a distinct carpophagus pit, a small but pointed pretarsus of the second maxillae, and 3-5 coxal pores in each coxopleuron with no isolated coxal pores.
Geophilus admarinus is a species of soil centipede in the family Geophilidae. It's found in southeast Alaska under stones near the low tide mark and is capable of surviving prolonged submersion underwater.
Geophilus alaskanus is a species of soil centipede in the family Geophilidae found in Alaska. It bears similarities to Mecistocephalus attenuatus, however unlike M. attenuatus, it has oblong cephalic lamina, unarmed coxae, and unarmed claw at base. G. alaskanus is dull orange-brown in color, grows to about 30 millimeters, and has 53 leg pairs with the first pair being very small and the last pair being distinctly larger than the others. It also possesses coalesced frontal lamina, unexposed basal lamina broader than the cephalic, a prehensorial sternum that's broader than it is long, distinct lateral grooves, obsolete prosternal teeth, a smooth, bare dorsal surface except for two faint impressed lines, and a small pleurae of the last segment marked with 8–10 pigmented pores of which the posterior one of the ventral face is larger than the others and sunk in a deep cavity.
Geophilus mordax, also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 millimeters in length, though it averages 25–40, has 49–53 leg pairs in males and 49–57 in females, and is bright red in color. G. mordax also bears 3–5 labral teeth, a short and robust apical claw of the second maxillae, an exposed prebasal plate, and a large, consolidated sacculi.
Geophilus nanus is a species of soil centipede in the family Geophilidae found on the Iberian Peninsula. It's a poorly defined species described as being pale yellow, 15 mm long, with smooth sternites bearing no furrows, final hip with few pores next to the sternite rim, tergites deeply double-furrowed from the basal shield onward, and 41 leg pairs. It’s apparently related to G. gracilis, differentiated by jaws that do not surpass the forehead or bear chitin lines, and by the 2nd tarsal segment of the final leg pair being just a tiny stub.
Diphyonyx conjungens, formerly Geophilus conjungens, is a species of soil centipede in the family Geophilidae that has been recorded in the Balkan Peninsula, Anatolia, the southern Sporades islands, the most eastern parts of Turkish Armenia, the Pontic and Tauric mountains, and Crimea. It grows up to 60 millimeters long and bears 69–81 leg pairs, as well as 1–2 stout tubercles and 2–4 slender filaments on the mid-part of the labrum, an absence of condyles between the anterior trunk sterna, and a single, isolated pore on each coxo-pleuron.
Geophilus richardi is a species of soil centipede in the family Geophilidae found in France, Italy, and the Ionian Islands. Females of this species have 33 pairs of legs; males have 29 or 31. This species is one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs. This species grows up to 10 millimeters long, has no carpophagus pit or pore-fields, and has a gradually tapering, curved pretarsus of the second maxillae. G. richardi lacks typical ventral pores between 2–4mm. The sternites instead bear a small number of pores between 0.5–1mm that differ from micropores, which are unbounded by a cuticular ring. These are possibly the remnants of typical ventral pores, their smaller size being a byproduct of overall miniaturization.
Geophilus winnetui is a species of soil centipede in the family Geophilidae found in Iowa. It grows up to 48 millimeters long and has 55–63 leg pairs, a well-developed labrum with the middle part having short, strong teeth, pushed backwards by median side parts that almost meet; maxilla with 2 pairs of somewhat blunt external palpi; coxal process not separated, with a number of strong bristles; and presternites undivided and so long that successive sternites are separated.
Geophilus bipartitus is a species of soil centipede in the family Geophilidae found in Japan. It grows up to 15 millimeters in length; the males have about 35 leg pairs, the females 39. It lives in Japanese white birch.
Geophilus rhomboideus is a species of soil centipede in the family Geophilidae found in Japan. It grows up to 30 millimeters in length; the males have about 43 leg pairs, the females 49.
Geophilus sounkyoensis is a species of soil centipede in the family Geophilidae found in Japan. Its body is yellow in color and it grows up to 40 millimeters in length; the males have about 55 leg pairs, the females 57.
Geophilus monoporus is a species of soil centipede in the family Geophilidae found in Tiba, Japan. It grows up to 18 millimeters in length; it's named for the single pore at the base of the final leg pair.
Geophilus dentatus is a species of soil centipede in the family Geophilidae found in Hokkaido. It's yellow in color, 25 millimeters in length, with 41-47 leg pairs and two clearly visible terminal pores. It's very similar to G. truncorum, but differs in the number of outer palpi on the 1st maxillae, the presence of denticles in the midpiece of the labrum, and the number of terminal pores.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 37 to 41 pairs of legs; females have 35 to 41.
Geophilus bosniensis is a species of soil centipede in the family Geophilidae endemic to Bosnia and Herzegovina. It grows up to 30 millimeters and has 75 leg pairs, as well as sternites unseparated in the median but with a suture line, and sternal pore areas in the trunk segments only. Overall, the identity and phyletic position of this centipede are uncertain.
