The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae. Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.
Geophilus is a genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus. Two species in this genus include centipedes with only 29 pairs of legs, the lowest number found in the family Geophilidae: G. persephones and G. richardi.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 40 millimeters, with between 65-73 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus admarinus is a species of soil centipede in the family Geophilidae. It's found in southeast Alaska under stones near the low tide mark and is capable of surviving prolonged submersion underwater.
Geophilus impressus is a species of soil centipede in the family Geophilidae found all over Europe, and has also been recorded in North Africa. It lives frequently in endogean habitats; in Sardinia it's found mostly in Quercus ilex woods, but also in Mediterranean shrub, open habitats, and maquis. It lives anywhere from sea level to 1700 meters above it, sometimes in caves.
Geophilus aetnensis is a species of soil centipede in the family Geophilidae found in Europe and northern Asia, excluding China. As described by Verhoeff in 1928, it grows up to 28 millimeters and has 53 leg pairs, 4 sensory setae each on the 2nd-4th front sternites, slightly notched maxillae, and very faint sternal pits on the thorax as well as poorly developed sternal grooves.
Geophilus gracilis is a species of soil centipede in the family Geophilidae found most commonly in Britain and Ireland, though specimens have also been recorded in Chile, France, Greece, and Algeria. It lives under mud and stones along the coast near or below the high tide mark, grows up to 30 millimeters in length, has between 51-57 leg pairs, and is bright yellow in color with a darker reddish head. It is often confused with G. flavus and, in coastal locations, G. osquidatum.
Geophilus mordax, also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 millimeters in length, though it averages 25–40, has 49–53 leg pairs in males and 49–57 in females, and is bright red in color. G. mordax also bears 3–5 labral teeth, a short and robust apical claw of the second maxillae, an exposed prebasal plate, and a large, consolidated sacculi.
Geophilus vittatus, also known as the diamondback soil centipede, is a species of soil centipede in the family Geophilidae widespread in North America and named for the dark band of diamond-shaped markings across its back. G. vittatus grows up to 52 millimeters in length, though it averages 25–40, and ranges from brown- to orange-yellow in color, with 49–53 leg pairs and a lack of consolidated carpophagus structures. It can be found under any debris on the forest floor, but mostly under loose bark and occasionally near the sea.
Geophilus alzonis is a species of soil centipede in the family Geophilidae found in Monte Alzo near Tolosa in Spain, which it was named after. It's a poorly defined species that was described as being light yellow with a light chestnut brown head and 13-14 mm long, with small but numerous leg bristles, sternites bearing three longitudinal furrows, a carpophagus formation from segments 3-6 to segments 11th-13, and 37-51 leg pairs.
Geophilus angustatus is a species of soil centipede in the family Geophilidae found on the Aleutian Islands. It's dark red, with 41–43 leg pairs; the antennae are cylindrical and about twice as long as the feet. Like other geophilomorphs, the antennae are 14-segmented. Its name comes from Latin 'angustatum', meaning 'narrowed', referring to its anteriorly narrowed body.
Geophilus anonyx is a species of soil centipede in the family Geophilidae found in Oregon. It was originally placed in the genus Brachygeophilus based on the lack of sternal pores, a character shared with the type species B. truncorum, however it was later moved to Geophilus.
Geophilus atopodon is a species of soil centipede in the family Geophilidae found in Beulah, New Mexico. Its body is uniformly light brown, up to 25 millimeters long and rather thick, narrowed posteriorly, with 49–51 leg pairs, a discrete frontal plate, and a long claw of the anal legs.
Geophilus richardi is a species of soil centipede in the family Geophilidae found in France, Italy, and the Ionian Islands. This species grows up to 10 millimeters long, has no carpophagus pit or pore-fields, and has a gradually tapering, curved pretarsus of the second maxillae. Females of this species have 33 pairs of legs; males have 29 or 31. G. richardi lacks typical ventral pores between 2–4mm. The sternites instead bear a small number of pores between 0.5–1mm that differ from micropores, which are unbounded by a cuticular ring. These are possibly the remnants of typical ventral pores, their smaller size being a byproduct of overall miniaturization. This species is one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs.
Geophilus winnetui is a species of soil centipede in the family Geophilidae found in Iowa. It grows up to 48 millimeters long and has 55-63 leg pairs, a well-developed labrum with the middle part having short, strong teeth, pushed backwards by median side parts that almost meet; maxilla with 2 pairs of somewhat blunt external palpi; coxal process not separated, with a number of strong bristles; and presternites undivided and so long that successive sternites are separated.
Geophilus monoporus is a species of soil centipede in the family Geophilidae found in Tiba, Japan. It grows up to 18 millimeters in length; it's named for the single pore at the base of the final leg pair.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 37 or 39 pairs of legs; females have 39 or 41.
Geophilus punicus is a species of soil centipede in the family Geophilidae found in the Western Mediterranean, specifically Italy and North Africa (Tunisia). It's an epigeic species, though its habitat preferences are unknown apart from one record from caves, indicating it may also be a trogloxene.
Geophilus bosniensis is a species of soil centipede in the family Geophilidae endemic to Bosnia and Herzegovina. It grows up to 30 millimeters and has 75 leg pairs, as well as sternites unseparated in the median but with a suture line, and sternal pore areas in the trunk segments only. Overall, the identity and phyletic position of this centipede are uncertain.
Geophilus claremontus is a species of soil centipede in the family Geophilidae found in Claremont, California, after which it was named. It was incorrectly placed in the genus Brachygeophilus in 1929 by Attems, most likely based on the lack of sternal pores.
