The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. Centipedes in this genus range from 1 cm to 8 cm in length. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Geophilus electricus is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named Scolopendra electrica in 1758, it was later moved to its own genus, Geophilus. As the first species described in this genus, it is considered by some to be the type species. It is yellowish-orange in color and can grow up to 45 millimeters, with 55 to 75 leg pairs, a distinct carpophagus fossae on the anterior sternites, and a distinctive arrangement of the coxal pores of the last leg pair. Like many geophilomorphans, G. electricus has been known to glow in the dark, most likely as a defensive maneuver.
Geophilus arenarius is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. This species is frequently misidentified with G. electricus, and as part of the carpophagus species-complex, is closely related to both G. carpophagus and G. easoni, though it differs mainly by lacking a transverse suture on the head and peculiar integumental features (carpophagus-structures) along the trunk, as well as having relatively stouter antennae and forcipular coxosternite. G. arenarius is distinctly smaller at full growth than G. carpophagus, with usually blunter and more sclerotised tubercles lining the intermediate part of the labrum and a minute denticle at the basis of the forcipular tarsungula. It has fewer bristles lining the lateral parts of the labrum than G. easoni as well as a generally higher number of legs and a more greyish coloured trunk. The descriptions of 36 G. arenarius syntypes indicate that males of this species have 55 pairs of legs, whereas females have 55 to 59 leg pairs, with 57 as the most common number.
Geophilus aenariensis is a species of soil centipede in the family Geophilidae found in the southern Italian Peninsula. It has a body length of up to 22 millimeters, around 47 leg pairs in females and 51 in males, an absence of anal pores and bristles on the lateral part of the labrum, a distinct carpophagus pit, a small but pointed pretarsus of the second maxillae, and 3-5 coxal pores in each coxopleuron with no isolated coxal pores.
Geophilus impressus is a species of soil centipede in the family Geophilidae found all over Europe, and has also been recorded in North Africa. It lives frequently in endogean habitats; in Sardinia it's found mostly in Quercus ilex woods, but also in Mediterranean shrub, open habitats, and maquis. It lives anywhere from sea level to 1700 meters above it, sometimes in caves.
Geophilus gavoyi is a species of soil centipede in the family Geophilidae. It is closely related to both Geophilus aetnensis and its junior synonym G. evisensis; it's considered by some to be a synonym of G. evisensis. It's found under stones in northern Asia and western Europe, especially France and England. It grows to between 23 and 35 millimeters in length and has between 39 and 43 leg pairs in males and 41–45 in females. In the since debunked elongata subspecies it was said to vary from between 49 and 55 leg pairs in males and between 51 and 57 in females. As described in 1964, it has well-developed sternal grooves, longer legs than G. aetnensis, and normal claws in the anal legs.
Geophilus aetnensis is a species of soil centipede in the family Geophilidae found in Europe and northern Asia, excluding China. As described by Verhoeff in 1928, it grows up to 28 millimeters and has 53 leg pairs, 4 sensory setae each on the 2nd-4th front sternites, slightly notched maxillae, and very faint sternal pits on the thorax as well as poorly developed sternal grooves.
Geophilus osquidatum is a species of soil centipede in the family Geophilidae found in western Europe, from mainland Spain through western France to Britain and Ireland, though it's also been recorded in Italy, Czech Republic, and Germany. Males of this species have 51 to 57 pairs of legs; females have 53 to 63 leg pairs. This species grows up to 30 millimeters and is bright yellow with a darker reddish head. Because of this, it's often confused with G. flavus and G. gracilis. Its subspecies, G. osquidatum porosum, was found synonymous with G. flavus. In Britain, G. osquidatum is found in a wide range of habitats including woodland, grassland, and coastal shingle as well as gardens and waste ground.
Geophilus mordax, also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 millimeters in length, though it averages 25–40, has 49–53 leg pairs in males and 49–57 in females, and is bright red in color. G. mordax also bears 3–5 labral teeth, a short and robust apical claw of the second maxillae, an exposed prebasal plate, and a large, consolidated sacculi.
Geophilus cayugae is a species of soil centipede in the family Geophilidae found in New York, Virginia, and North Carolina. It grows up to 68 millimeters in length, though it averages 25–40, has 57–65 leg pairs in males and 63–67 in females, and is bright whitish yellow in color. G. cayugae can be identified by its well-developed paxilli, weakly consolidated sacculi, and concealed prebasal plate combined with lack of consolidated prebasal areas. It's found beneath rocks and other debris in high elevation montane regions.
Geophilus vittatus, also known as the diamondback soil centipede, is a species of soil centipede in the family Geophilidae widespread in North America and named for the dark band of diamond-shaped markings across its back. G. vittatus grows up to 52 millimeters in length, though it averages 25–40, and ranges from brown- to orange-yellow in color, with 49–53 leg pairs in females, 49–51 leg pairs in males, and a lack of consolidated carpophagus structures. It can be found under any debris on the forest floor, but mostly under loose bark and occasionally near the sea.
Geophilus alzonis is a species of soil centipede in the family Geophilidae found in Monte Alzo near Tolosa in Spain, which it was named after. It's a poorly defined species that was described as being light yellow with a light chestnut brown head and 13–14 mm long, with small but numerous leg bristles, sternites bearing three longitudinal furrows, a carpophagus formation from segments 3–6 to segments 11th-13, and 37-51 leg pairs.
Geophilus anonyx is a species of soil centipede in the family Geophilidae found in Oregon. It was originally placed in the genus Brachygeophilus based on the lack of sternal pores, a character shared with the type species B. truncorum, however it was later moved to Geophilus.
Geophilus truncorum is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a yellow or orangeish brown body and dark yellow or brown head, denser and shorter hair than most Geophilus species, a main plate almost as elongated as in G. flavus (115:100), and distinct carpophagus fossae on the anterior sternites. Males of this species have 35 to 41 pairs of legs; females have 37 to 41.
Geophilus pusillus is a species of soil centipede in the family Geophilidae found in Algeria. It grows up to 11 millimeters in length. The original description of this species is based on ten specimens: five male specimens from Algeria with 31 pairs of legs, and five specimens from Germany. Records from the Alpstein mountains indicate that G. pusillus is a soil-dwelling species that prefers humus-rich soil, but these records deserve confirmation. Some authorities consider the specimens recorded from Europe to be specimens of G. ribauti that have been misidentified as specimens of the Algerian species G. pusillus.
Geophilus punicus is a species of soil centipede in the family Geophilidae found in the Western Mediterranean, specifically Italy and North Africa (Tunisia). It's an epigeic species, though its habitat preferences are unknown apart from one record from caves, indicating it may also be a trogloxene. The original description of this species is based on male specimens with 43 pairs of legs.
Geophilus bobolianus is a species of soil centipede in the family Geophilidae found in France and Italy. This species has 45 to 51 pairs of legs. It was originally classified as a subspecies of G. longicornis identified by its lack of anterior sternal pores.
Geophilus crenulatus is a species of soil centipede in the family Geophilidae found in a ravine above the Hemis monastery in India, 3525 meters above sea level. It was originally thought to be a subspecies of G. intermissus and named Geophilus intermissus var. crenulata. The original description of this species is based on a specimen measuring 22 mm in length with 57 pairs of legs.
Geophilus intermissus is a species of soil centipede in the family Geophilidae found in India around the Himalayas. The original description of this species is based on a female specimen measuring 35 mm in length, with 57 pairs of legs, a yellowish red anterior, and especially thin tarsi on the last leg pair.
